188 



Anaplecta 



In the previous species LI and pne are in the central dorsal wall. In Anaplecta the 

 corresponding area is membranous and just somewhat depressed (fig. 209). This area is 

 interpreted as the vestige of a pne-pouch, with LI completely lost. This assumption is 

 supported by the muscles 12 and 13 (fig.201, 221), which run to L4-sclerotisations in the 

 left edge of the left complex (12) and to L2 in the dorsal wall of the Ive-pouch (13). These 

 are the same connections as in the species discussed before. 11 is, like in some other 

 species, missing. (The homology of L4 and L2 is discussed in 6.2.4. and 6.3.4). Further 

 support comes from McKittrick (1964): She identifies in another species of Anaplecta 

 ("sp. C) a sclerite LI (McKittrick's fig. 11 2), which has the same position as the assumed 

 pne-vestige in the species I studied. 



Nahublattella 



LI' and pne (fig. 242, 243) show some characteristic features: They take a dorsal position. 

 The level anterior part of LI' (Lla in fig.323n) occupies the ventral wall of a pouch pne. 

 The posterior part of LI' completely sclerotises a bulge-like process (dca in fig. 243, 244). 

 This is interpreted as a sclerite-ring (regions Lll, Llm, and Llr in fig.323n) encircling 

 the dea-process which has spread posteriad over the whole dca (similar to but more 

 complete than in Tryonicus parvus, compare fig.323g and n). Further arguments for 

 homology come from the muscles on pne: 11 (fig. 249) runs leftward to the sclerite-ribbon 

 L4d' (which is probably homologous with L4d of Mantoida, fig.44, and Cryptocercus, 

 fig. 150: discussion in 6.3.4.; compare 11 of Mantoida, fig.48, and Cryptocercus, fig. 155). 

 13 (fig.250) runs to L2 in the dorsal wall of the Ive-pouch (compare 13 of e.g. Mantoida, 

 fig. 50, Polyphaga, fig. 128, Cryptocercus, fig. 158, 159, and Anaplecta, fig.221); the 

 homology relations of L2 and Ive are discussed in 6.2.). 19a (fig. 249) runs to the membrane 

 to the right of LI' - as do 19 in Anaplecta (fig.221) and the posterior part of 19 in 

 Polyphaga (fig.127, 129). 



The right insertion of 12 (fig. 249) has shifted away from the pne-pouch (discussion in 

 6.3.4.). Muscle 128, with both insertions on LI', is pecuhar to Nahublattella (fig.250). 

 Like in Cryptocercus, but in contrast to the other species, LI and L2 are no longer in 

 contact (articulation A2 lost). 



Parcoblatta and Blaberus 



There are no vestiges of LI and pne. Muscles 11 and 13 have been lost. 12 has shifted in 

 the same way as in Nahublattella (discussion in 6.3.4.). 



6.2. Left complex II: Main sclerite L2 and associated elements 

 6.2.1. Comparison between Blattaria and Mantodea 



In the Mantodean and in several Blattarian species the L2-sclerotisations, the Ive-pouch, 

 the vla-lobe, the processes paa and pda, and the genital opening show the same principal 

 arrangement and similar positions relative to pne and LI. The proportions of these 



