191 



left parts of the vla-lobe and the Ive-pouch. In all three species the invagination of the 

 Ive-pouch (edge 7 in e.g. fig. 96-98) and the free left edge of the vla-lobe (edge 61 in e.g. 

 fig. 87, 98) start at the same point in the ventral wall of the left complex. Anterior to this 

 point the ventral vla-wall is confluent with the remaining ventral wall of the left complex 

 (e.g. fig. 87). This point takes a different position in the three species. In Tryonicus it is 

 far anteriorly (fig. 87): The left edge 61 of via extends far anteriad, and the invagination 

 of the Ive-pouch begins far anteriorly. In Mantoida this point is by far more posteriorly 

 (7 and 61 in fig.41, 47): The left edge 61 of via does not extend so far anteriad, the ventral 

 vla-wall is except for its posteriormost part confluent with the remaining ventral wall of 

 the left complex, and the invagination of the Ive-pouch begins far posteriorly. In Polyphaga 

 this point is at the posterior edge of the left complex (fig. 122, 123): The vla-lobe does 

 not have a free left edge 61 at all, the ventral vla-wall is completely confluent with the 

 remaining ventral wall of the left complex (fig. 11 5), and the invagination of the Ive-pouch 

 begins most posteriorly. The ground-plan state of this feature is assumed to be somewhere 

 in between the situation of Mantoida and that of Polyphaga: 



17. The ventral vla-wall is for most or all of its length confluent with the remaining ventral 

 wall of the left complex, the left edge 61 of the vla-lobe ends far posteriorly or is 

 missing, and the invagination of the left(-ventral) part of the Ive-pouch begins far 

 posteriorly. 



Archiblatta likewise has a second pouch (Ive in fig. 54, 55) in the right part of the left 

 complex. Like in Polyphaga and Mantoida, the sclerotisation L2 (fig. 55) runs hke an arch 

 along the edges of the pouch (7 in fig. 55). To the same extent as in Tryonicus, the right 

 parts of L2 and Ive curve dorsad and back to the left, and in this area L2 is very broad. 

 Similar to Mantoida (fig.46, 47), L2 is mostly restricted to the inner = dorsal Ive-wall but 

 bends into the outer = ventral Ive-wall in the posterior left part of Ive (at the posterior 

 end of edge 7 in fig. 55, 56). In some features Archiblatta coiTcsponds with all previous 

 species: L2 articulates with LI (A2 in fig. 54, compare fig. 94, 118, 45, 46). The left 

 posterior part of L2 leaves the Ive-pouch and runs onto a process (paa in fig. 55, 56), 

 which thus corresponds to the paa of the other species in this aspect of its relative position. 

 The outer = ventral Ive-wall is the dorsal vla-wall (fig.53; via is pulled to the right), is 

 membranous, and receives the ejaculatory duct (D in fig.53). The ventral vla-wall is a 

 right part of the ventral wall of the left complex and is largely sclerotised (by L4G in 

 fig. 54). Like in Polyphaga only, the vla-lobe does not have a left edge 61 but its ventral 

 wall is entirely confluent with the remaining ventral wall of the left complex. 

 Derived features of Archiblatta are (1) that paa is mostly membranous, (2) that there is 

 no connection between the sclerotisations of paa (L2) and pda (L4), and (3) that paa is 

 quite far removed from pda (compare fig.44 and 53). The two latter features are probably 

 correlated with a derived feature of the Ive-pouch: (4) The posteroventral part of Ive has 

 strongly receded to the right (compare edges 7 in fig. 122, 46, and 55) and is restricted to 

 a narrow right part of the left complex. The anteroventral part of Ive is still deeply 

 invaginated to the left and has become tongue-like. 



Eurycotis is similar to Archiblatta but in some features more derived (compare fig. 67, 68 

 and 54, 55): (1) L2 is not an arch but a plate (probably the arms of the arch have fused). 



