193 



6.2.2. The elements in the common ground-plan of Blattaria and Mantodea 



The features 1.-18. in 6.2.1. permit a reconstruction of the ground-plan moiphology of 

 L2, Ive, paa, and via, and of the genital opening (fig.321e,g): The Ive-pouch lies ventral 

 to the pne-pouch; it is quite broad but does not reach the left edge of the left complex 

 (and does not have a recess to the right in its posterior part). L2 is largely restricted to 

 the dorsal Ive-wall and runs like an arch along the lateral and anterior edges of the Ive- 

 pouch. The left part (L2p-region) of the arch leaves the pouch posteriorly, and the 

 posteriormost part (L2d-region) sclerotises the short, somewhat upcurved process paa. 

 L2d is connected with the sclerotisation of the process pda, and paa and pda are close 

 to each other. The right end of L2, or the dorsal left end in the case of L2-upcurving 

 (L2m-region), articulates with Llm (A2). The ventral Ive-wall is mostly membranous and 

 is at the same time an anterior part of the dorsal vla-wall. The ejaculatory duct D opens 

 into the right anterior part of this membrane. The ventral vla-wall is largely sclerotised 

 and almost completely confluent with the ventral wall of the remaining left complex (i.e. 

 the left edge 61 of via is missing or does not extend far anteriad). Muscles 13, 14, 15, 16a, 

 and 16b are present. 



6.2.3. Homology relations and character states of the elements in Mantodea 



In Chaeteessa, Metallyticus, and Sphodromands, the main features of L2, Ive, and paa 

 are like in Mantoida (fig. 11, 26, 34, 45, 46): Ive lies ventral to the pne-pouch. L2 is 

 restricted to the dorsal Ive-wall. The right parts of L2 and Ive are level. Posteriorly L2 

 leaves the Ive-pouch to provide the sclerotisation of a process which is therefore regarded 

 as paa. The right margin of L2 articulates with LI (A2; compare in 6.1.3.). 

 In contrast to Mantoida, Metallyticus and Sphodromantis have separated the sclerotisations 

 of paa (L2) and pda (L4), and the processes are more distinct from each other and more 

 prominent (fig. 10- 12, 23-26, 44-46). In Chaeteessa (fig.31, 32, 34) paa is very prominent, 

 too, but a process pda is missing. Nevertheless, in this species, too, L2 and L4 are 

 completely separated in the area concerned. 



L2 of Chaeteessa, Metallyticus, and Sphodromantis has become plate- or ribbon-like and 

 has spread over most of the dorsal Ive-wall. Like in Eurycotis, the arms of the arch (regions 



Fig.324: Left complex, homologous regions of main sclerite L2 (on pages 194, 195). - Only L2- 

 sclerotisations are shown. Dorsal views. L2 is divided into the four regions L2a, L2d, L2m, and 

 L2p (definition in 6.2.1.); the extensive parts of L2 in the ventral wall of pouch Ive in Polyphaga 

 are labelled L2v (not defined as a region of L2). If L2 is divided into several sclerites, these are 

 labelled with the capital letters used in the text and in fig. 1-3 19 (e.g. B = L2B). Undulate lines are 

 cutting lines through sclerotisations (if L2 is fused with parts of other main sclerites, e.g. L4). In 

 fig.324e,f,g the left drawing shows the complete L2, the right drawing shows L2 after removal of 

 its dorsal parts. The part of the sclerite margin which forms articulation A2 with sclerite LI is 

 indicated by dashes. A4 and AlO are articulations between L2-sclerites. paa and via are processes 

 occupied at least partly by L2-sclerotisations. The sclerite of Parcoblatta (fig.324n) and Blaberus 

 (fig.324o) which is termed L2 in the text has to be designated correcüy L2D+(L2E+L4N), compare 

 in 6.2.4.; the L4N-part has been removed from the sclerite. 



