197 



a split off part of the Llm-region like LIB of Sphodromantis, fig. 10?). This equals the 

 question which of the articulations A2 (fig. 178) and A4 (fig. 180) is the true A2. 



- In Mantoida, Polyphaga, and Tryonicus the left posterior part of L2 leaves the pouch 

 and continues into the sclerotisation of the processes paa and pda (fig.46, 96, 118). In 

 Lamproblatta, the left posterior part of L2A also has an extension that leaves the pouch 

 (L4S in fig. 178, 180), and right-anterior to this area there is a sclerotisation with two 

 processes (paa and pda in fig. 178, 179), which, however, is completely separated from 

 L2A and L4S). The question arises if paa and pda of Lamproblatta are homologous 

 with those of the other species. 



These problems can be solved by a comparison of the musculature of Lamproblatta and 

 the other species - especially Polyphaga. 



- Lamproblatta also has a stout muscle from LI to Ive (13 in fig. 187; compare Polyphaga, 

 13 in fig. 128). Its insertion area on Ive is within the recess and also includes parts of 

 L2B. 



- Lamproblatta and Polyphaga have 4 muscles (15, 16a, s3, sl2) inserting close to each 

 other in the anteriormost ventral wall of the left complex (fig. 133, 188); homology is 

 assumed for all of them (s3 and sl2 are discussed in 6.9.). In Polyphaga the two 

 posterior muscles (15 and 16a) run to L2 at the left or right, respectively, anterior edge 

 of the Ive-pouch. In Lamproblatta 15 runs to L2A on the left of the recess, 16a runs to 

 L2B on the right of the recess. 



- The two 16b-bundles (fig. 189) of Lamproblatta are assumed to be homologous with the 

 16b of the other species {Polyphaga: fig. 132): The ventral insertion is posterior to that 

 of 16a. The insertion in the dorsal vla-wall is somewhat different in Polyphaga and 

 Lamproblatta (next to the genital opening or far posterior to it), but the insertions of 

 the two bundles of Lamproblatta are similarly situated as the insertion of the one bundle 

 of 16b of Sphodromantis (fig. 18) or Cryptocercus (fig. 155, 157, see below). 



- Muscle 112 of Lamproblatta and Polyphaga inserts in the right part of the ventral = 

 outer Ive- wall - very close to the dorsal insertion of 16a (fig. 128, 129, 186, 188) - and 

 runs to a small sclerite in the right dorsal wall of the left complex (L8, homology 

 discussion in 6.5.). Homology is assumed for these 112. In Lamproblatta 112 inserts to 

 the right of the recess, on L2B. (L8 and 112 are derived features of Polyphaga, Ergaula, 

 and Lamproblatta.) 



- Muscle 110 of Polyphaga (fig. 129) runs from L2 in the left dorsal wall of Ive to the 

 sclerotisation in between the processes paa and pda. Lamproblatta has a muscle (110 

 in fig. 186) from the same area of Ive to the sclerotisation between paa and pda. Such 

 a muscle is missing in all species discussed before. 



The course of 110 suggests homology for paa and pda of Lamproblatta and Polyphaga. 

 Since the pda-sclerotisation is part of L4, the discussion of this topic will be continued 

 in 6.3.4.. 



The insertions of 13, 16a, and 112 clearly demonstrate that L2B is a true part of L2 

 (regioning in fig.324i) and that the respective part of the pouch is a true part of Ive. That 

 the right part of L2B curves dorsad and leftward like the right part of L2 in other Blattaria 

 and that the genital opening is to the right of the recess (fig. 122, 180) supports this 



