204 



of Anaplecta (fig. 222) also inserts on the ejaculatory duct but bridges the vfa-outfolding 

 on its way to its insertion on the dorsal base of via (discussion of 113 in 6.5.). Muscle 

 16a of Anaplecta (fig. 222) resembles 16a of Polyphaga and Lamproblatta (fig. 133, 188) 

 in inserting ventrally behind s3 and dorsally at the right anterior edge of the Ive-pouch. 

 Whether muscle 15 is homologous with 15 of the other species (fig. 133, 188, 223) is 

 questionable: The insertion on L2 is similar in Anaplecta and e.g. Polyphaga; the anterior 

 insertion (on L4), however, is situated quite differenUy in these two species. Homology 

 is also unclear for the muscles 125 and 126 (fig.224). 



As compared with the previous species, Anaplecta has some important derived features: 

 (1) The anterior part of the Ive-pouch is a tube-like Ive-apodeme. (2) Edge 7 is interrupted 

 by vfa and vpe. (3) The common sclerotisation of paa and pda is stout and ring-shaped 

 at its base. (4) Muscle 14 from L2 to left parts of L4 has been lost (like in Lamproblatta). 



Nahublattella 



The part of the left complex comprising the large pouch Ive (fig. 242), the opening of the 

 ejaculatory duct (D in fig. 242), the processes, via, vsa, paa, and pda (fig. 244, 245), and 

 the sclerotisations L2D', L2E', and L4N' show a lot of similarities with the elements of 

 Anaplecta discussed before: 



All these elements lie in the center of the left complex and (antero-)ventral to the pne- 

 pouch. The anterior part of the Ive-pouch (see edges 7 in fig. 242) is a tube-like Ive- 

 apodeme, whose dorsal wall is completely sclerotised, and whose ventral wall contains a 

 membranous stripe (44 in fig. 206, 212, 239a, 245). Muscle s7 runs from the Ive-apodeme 

 to the subgenital plate (fig.200, 234). 



At the base of the apodeme, L2D' is somewhat forked (fig. 243, 324m), like L2 in 

 Anaplecta (fig. 2 12, 3241): Extension 36 is the left branch, the posterior main part of L2D' 

 is the right one. At the left branch there adjoins a ring-shaped sclerotisation lying at the 

 base of some processes (paa and pda in Anaplecta, fig.2 11-214; via with vsa, paa, and 

 pda in Nahublattella, fig.244, 245). A stout muscle 110 (fig.222, 250) runs from the Ive- 

 apodeme to the left part of this sclerite-ring. However, in Nahublattella the sclerite ring 

 has become separated from the rest of L2 (L2D') by an articulation (AID in fig.244). At 

 the base of the left branch the L2-sclerotisation bends into the ventral Ive-wall and forms 

 a posterior extension (28 in fig.2 15, 245). However, in Anaplecta the cuticle around 

 extension 28 forms a process (gta in fig.2 15), which is missing in Nahublattella. 

 The right branch of L2 or L2D', respectively, extends rightward in Anaplecta but more 

 posteriad in Nahublattella. The relation between this sclerotisation and the dorsal wall of 

 the ejaculatory duct (D in fig.245, 246) is the same in the two species (fig.211, 245), but 

 only in Nahublattella the right-anterior margin of this L2-part folds narrowly back to the 

 left (towards edge 38 in fig.245). 



Two further muscles inserting on L2D' of Nahublattella correspond with muscles of the 

 other species: 13 running to the pne-pouch (fig. 250, compare e.g. fig. 50, 71, 128, 221), 

 and 14 running to L4-sclerotisations in the left part of the left complex (fig.249, compare 

 e.g. fig.50, 71, 129; missing in Anaplecta; homology discussion of L4 in 6.3.4.). 



