206 



The membranous lobe via (fig. 239a, 245-247) has similar features as the vla-lobe of e.g. 

 Sphodromantis (fig.6,12), Lamproblatta (fig. 174, 180), Cryptocercus (fig. 148, 151), and 

 Eurycotis (fig. 63, 66): Its ventral wall is part of the ventral wall of the left complex. Its 

 dorsal and its ventral walls are connected by a stout muscle 16b (fig.251, 252, 18, 71, 155, 

 188, 189). (These two features are also true of Anaplecta, fig.205, 218, 224). Its dorsal 

 wall is at the same time the ventral wall of the Ive-pouch. (This is not true of Anaplecta 

 because of the membrane foldings between Ive and via, especially vfa). 

 Like the other species (with the exception of Cryptocercus), Nahublattella has muscles 

 from the anterior ventral wall of the left complex to the L2-sclerotisations: 15 is certainly 

 homologous with 15 of Anaplecta (similar posterior insertion on the left branch of L2; the 

 homology of the anterior insertion is discussed in 6.3.4.), but, as in Anaplecta, homology 

 with the 15 of the other species is questionable. Muscle 16a (fig. 250) could be homologous 

 with 16a of Anaplecta (and the other species); however, the insertion on the Ive-apodeme 

 is by far more anteriorly (fig. 222, 250), and the insertion in the ventral wall is on 

 sclerotisation. Alternatively, homology with 126 (fig. 224) of Anaplecta seems possible. 

 Nahublattella shows some important derived features as compared with Anaplecta and, at 

 least in the case of (2)-(6), all other previously discussed species: (1) The right branch of 

 L2 (posterior part of L2D') is by far narrower (compare fig. 213 and 245). Moreover, the 

 whole right posterior dorsal part of the left complex - that part with the right L2-branch 

 in its ventral wall {Anaplecta: fig.211-213) - is strongly reduced to form just the bifid 

 psa-process (fig. 245; compare fig. 328a and b). (2) L2 has divided into L2D' and L2E' 

 by articulation A 10. (3) The sclerotisation at the common base of the processes paa and 

 pda, which is ring-shaped in Anaplecta (fig.211-213), has lengthened to form a cylinder 

 (fig. 244). Hence, the processes paa and pda (and vsa) are now only the distal branches 

 of a larger evagination, which has been defined as a "new" process via (paa, pda, and 

 vsa are subordinate parts of via). The homologies of the single processes of Anaplecta 

 and Nahublattella are hardly determinable, but in my view the relations expressed by the 

 designations are the most probable. In accordance with Anaplecta, the sclerotisation of 

 via is assumed to comprise a L2-part (L2E', roughly the L2d'-region; dorsally on via 

 and near articulation AID; fig.3241,m) and L4N (ventrally on via and near the insertion 

 of 110; fig.3251,m; discussion of L4N in 6.3.4.). (4) This sclerotisation of via is divided 

 into a basal and a distal sclerite (39 in fig.241, 244). (5) There is no sclerotisation in the 

 ventral wall of the vla-lobe (compare fig.205 and 239a). (6) The main muscle of the hla- 

 hook (114 in fig. 249) has its anterior insertion on the Ive-apodeme (discussion in 6.4.). 



Parcoblatta, Blaberus, and other Blattelhdae and Blaberidae 



The L2-sclerotisations, the pouch Ive, the processes via, paa, pda, and psa, the lobe via, 

 and the muscles s7, 14, and 110 have been studied not only in Parcoblatta and Blaberus 

 but also in Supella, Euphyllodromia, Loboptera, Ectobius and Nyctibora (Blattelhdae), 

 Nauphoeta and Blaptica (Blaberidae) (muscles not studied in Ectobius). Morphology and 

 homology of these elements are shown in fig. 328. The morphology of all these species is 

 derived from a situation similar to Nahublattella. 



