211 



to the left base of via (110 in fig.250, 328b,h). Thus, the bipartition of L2 (by AlO: L2D 

 and L2E) is assumed to be homologous in Nyctibora and Nahublattella, and for via of 

 Nyctibora the same composition of L2E and L4N is assumed as for via of Nahublattella 

 (further details on L4N in 6.3.4.). In some features Nyctibora is more derived than 

 Nahublattella: (1) The right posterior branch of L2D (fig. 244, 245; compare fig. 328b and 

 h) is reduced to a vestige, and the process psa is completely missing. (In Nyctibora the 

 vestige can be identified by the insertion of 14, which is much closer to articulation AlO 

 than in Nahublattella: fig.328b,h). (2) The posterior insertion of 110 is upon a long 

 cuticular tendon (tve in fig.328h). (3) The via-process is no longer forked, and paa and 

 pda (and vsa?) must be fused or partly reduced. 



Nauphoeta (fig.328i) strongly resembles Nyctibora: The via-process, the articulation AlO, 

 the insertions of 14 and 110, the tve-tendon, and the phallomere-gland are arranged in the 

 same way (compare fig.328h and i). However, the basal sclerotisation of via is no longer 

 a complete cylinder and does not reach the base of the tve-tendon. 

 Supella (fig. 328c) is similar to Nauphoeta, but some features are different: The phallomere- 

 gland and the tve-tendon are missing. The sclerotisation of via has expanded anteriad 

 along the right margin of L2D', and articulation AlO is therefore long and hinge-like. The 

 right insertion of 14 is in the usual position but has shifted from L2D' to the adjacent 

 membrane. The posterior insertion of 110 has shifted to the right; its position can be 

 explained by a clockwise (as seen from behind) rotation of the via-process along its 

 longitudinal axis (similar to Parcoblatta, see below). The anterior UO-insertion on L2D' 

 is by far more posteriorly than in the other Blattelhdae and Blaberidae having a 110; 

 however, in Anaplecta, fig. 222, and Nahublattella, fig.250, the UO-insertion also extends 

 far posteriad. 



Alternatively, one could assume that in Supella the process is not via but psa (compare 

 fig. 328b and c) and that via is missing. However, no muscle in any of the Dictyopteran 

 species studied here would then have the same course as 110 of Supella (from the anterior 

 part of L2 to its right part), and articulation AlO of Supella would likewise have no 

 homologue at least in Blattellidae and Blaberidae. Therefore, and since the respective area 

 is quite similar in Supella and Nauphoeta, the process is more likely to be via. 

 Euphyllodromia (fig.328d), Parcoblatta (fig.328e), and Blaberus (fig. 328k) have, in 

 contrast to Nahublattella, Nyctibora, Nauphoeta, and Supella, the sclerotisation of via 

 firmly connected with the L2-sclerotisation of the Ive-pouch (like in Anaplecta and in the 

 ground-plan), and muscle 110 is missing (however, 142 of Blaberus, fig. 304, might possibly 

 be a 110 with its posterior insertion shifted far to the left). The basal sclerotisation of via 

 is a complete cylinder (fig.328d, 273, 274, 300, 302). Muscle 14 is present (fig.328d,e,k). 

 In Parcoblatta the via-process and the surrounding area have undergone a rotation 

 (clockwise as seen from behind; lower curved arrow in fig.328e). This can be recognised 

 by the following features: (1) The contact between the lumina of the via-process and of 

 the rest of the left complex (fig.328e, 273, 274) is dorsal to the connection of the 

 sclerotisations of via and Ive, not to the left of this connection as e.g. in Nyctibora 

 (fig.328h) and Nahublattella (fig. 328b). (2) The right part of the Ive-pouch, including the 

 distal part of the ejaculatory duct, has partly wrapped around the L2-sclerite (again. 



