216 



and L41 (? in fig.325d). Hence, 9. is an uncertain ground-plan feature. In Mantoida there 

 is no indication for the presence of a L4n-region, and a nla-process is missing. 

 Evidence from Chaeteessa (complete discussion in 6.3.3.) suggests that the L4n-region is 

 also an element of the common ground-plan of Blattaria and Mantodea and that the lack 

 of L4n in Mantoida is derived: 



11. The heavier sclerotised transverse bridge in the anterior ventral wall of the left 

 complex of Chaeteessa might, according to its very similar position, well be 

 homologous with the L4n-region of Eurycotis (compare fig. 31 and 65, 69; fig.325c,e). 

 Tryonicus has some features in common with Archiblatta and Mantoida which suggest the 

 homology relations shown in fig.325d,f,g. The two sclerites L4K and L4N (fig. 85, 97) 

 together form a broad ribbon in the left wall of the left complex, which takes, hke L4C 

 in Archiblatta (fig. 54), a position left-dorsal to the base of the hla-hook. Most of L4K 

 and L4N is therefore assumed to represent the L41-region. The nla-process (fig.97) on 

 L4K corresponds to the nla of Archiblatta (fig. 56) and Eurycotis (fig.68) in its shape, in 

 its location in the anterior ventral wall, and in its position relative to the other L4- 

 sclerotisations and to the hla-base. Thus, the part of L4K on nla is regarded as the L4n- 

 region. That part of L4K which anterior to nla extends to the right (fig.95) has the same 

 relative position as the right-anterior part of the L4C-crescent of Archiblatta (fig. 53, 55) 

 and is hence assumed to belong to the L41-region. The ribbon-like extension of L4N (L4d 

 in fig. 88-95) corresponds in its shape and relative position with the L4d-region of 

 Mantoida (fig.45) and especially Archiblatta (fig. 53). The posteriormost part of L4N 

 occupies a process (pda in fig.91) like in Mantoida and Archiblatta (pda in fig.44, 53) 

 and can be regarded as part of the L41-region. Like in Mantoida but in contrast to 



Fig. 329: Left complex, subdivisions of regions L41 and L4d into individual sclerites in Blattaria and 

 Mantodea. - Region L4I (discussion in 6.3.) is in its primitive condition undivided and connected 

 with region L2d posteriorly (between processes paa and pda). In many species L4I is connected 

 with region L4n anteriorly (ground-plan condition unclear). The various subdivisions of L41 in the 

 subgroups of Blattaria and Mantodea and the hypothetic directions of evolutionary transformation are 

 shown. The various types - or further derivations of them - are present in the species listed. It is 

 intended to emphasise principal similarities and differences in the subdivision of L41. For 

 comparability, all elements are left in their most primitive condition (like in fig.329a) - except for 

 the subdivisions of L41 and of the included parts of L2. Inner views from the right side; dorsal«—, 

 ventral-^, anteriort, posteriori. 

 The following structures are shown: 



- The sclerite regions L41 (white) and L4d (light pattern) and parts of L2 (dark pattern; mainly 

 region L2d on process paa). The right-anterior undulate line (1 in fig. 329a) represents the removal 

 of region L4n. The right-posterior undulate line (2 in fig. 329a) represents the removal of the 

 remainder of L2. L4d is always shown in its most primitive position, orientation, and shape, even 

 if these have changed or if L4d has been lost. 



- The dividing lines which cause a division of these sclerotisations into individual sclerites. Dividing 

 lines along which the respective sclerites are sdll in close contact are labelled A+Number 

 (articulation, e.g. A5; like in the text); if the respective sclerites are farther away from each other 

 the name of the respective articulation is put in brackets (e.g. (A5)). 



