223 



from the pne-pouch to the left and dorsad (homology discussion in 6.1.1.)- The insertion 

 of 11 is in Cryptocercus on L4N (including L4d), in Mantoida on and to the right of 

 L4d. 



- In Cryptocercus and Mantoida, muscle 19 (fig.49, 155) runs transversely within the 

 dorsal wall of the left complex (homology discussion in 6.5.). Its left insertion is in 

 Cryptocercus anterior to the left end of the assumed L4d-part of L4N, in Mantoida near 

 the right end of L4d; regarding the orientation of L4N assumed for Cryptocercus, these 

 ends of the sclerites would be homologous. In Cryptocercus, however, 19 does not extend 

 as far to the right as in Mantoida. 



Some features of Cryptocercus are derived: (1) The sclerotisation of pda is largely reduced 

 (fig. 150, 329h), and, possibly as a consequence of this, (2) the sclerotisations of pda (L4) 

 and paa (L2) have separated (like in Chaeteessa, Metallyticus, Sphodromantis, Archiblatta, 

 and Eurycotis). 



L4K-sclerite, process nla L4K of Anaplecta (fig. 209, 212) is probably 

 homologous with L4K of Tryonicus (fig. 98); similarities are: 



- The position in the left and left-ventral walls of the left complex. 



- The anterior part is on a bulge-hke evagination (nla in fig. 97, 212). Like in Tryonicus, 

 this sclerotisation is regarded as the L4n-region (fig.325g,l). 



- The posterior part is plate-like, with a broadly truncate posterior margin, and this part 

 partially encloses the retracted hla-hook from the left side (fig. 85, 202). Like in 

 Tryonicus, this sclerotisation is regarded as an anterior part of the L41-region 

 (fig.325g,l). 



A comparison of the muscles of Anaplecta and Eurycotis confirms these assignments to 

 L4n and L41: 



- That in both species the anterior insertion of the main muscle of the hla-hook (114c or 

 114; fig. 73, 222) is on or near the sclerotisation of the nla-bulge shows the homology 

 of these nla. The nla-sclerotisation of Eurycotis is, by definition, the L4n-region 

 (fig.325e,l). 



- In Eurycotis and Mantoida the muscles 12 (fig.49, 70) and 14 (fig. 50, 71) insert close 

 to each other on the L41-region and run to the pouches pne and Ive, respectively. 

 Anaplecta also has a muscle 12 from the posterior part of L4K to the pne-"pouch" 

 (fig.221, compare in 6.1.4.); muscle 14 is missing. Nahublattella (complete discussion 

 below), however, has both 12 and 14 (to the pouches pne and Ive), and their left insertions 

 are on a sclerotisation homologous with the posterior part of L4K of Anaplecta (L4U' 

 in fig.249). 



The narrow sclerotisation which extends in Tryonicus from the anteriormost part of L4K 

 to the right (anterior to nla, fig.96, 97, 325g) is missing in Anaplecta (fig.3251). Since 

 this sclerotisation probably corresponds to the anteriormost L41-region of Archiblatta and 

 Mantoida (bearing the anterior part of the swe-apodeme; fig.44, 53, 325d,f), this feature 

 of Anaplecta is regarded as derived. 



The course of the dividing line through the L41-region of Anaplecta (separating L4K and 

 L4N) can be deduced from the positions of the 12- and 14-insertions in Eurycotis, 



