224 



Mantoida, Anaplecta. and Nahiiblattella and from the distribution of the other 

 substructures: Anterior to the line (on L4K) there are the insertion of 12 (and 14 in 

 Nahiibkmella) - and hence parts of the L41-region - the L4n-region, the nla-process, and 

 the 114-inseilion. Posterior to the Hne (on L4N) are the pda-process - and hence posterior 

 parts of the L41-region - and the UO-insertion (and the L4d'-extension in Nahiiblattella). 

 The course of the line is shown in fig.329e. The distribution of all cuticular elements 

 present is the same as in L4K and L4N of Tn'oniciis. and the dividing lines of Anaplecta 

 and Tiyoniciis (and Nahiiblattella) are strongly suggested to be homologous. 

 L4K of Ctyptocerciis (fig. 150, 151) is homologous with L4K of Tiyoniciis and Anaplecta 

 and is likewise composed of the anterior L41-region and of the L4n-region. Both L41 and 

 L4n. howe\'er. are strongly reduced. These relations are, firstly, suggested by similarities 

 in the cuticular elements of Tiyonicus and Cnptocercus: 



- L4K of Ciyptocercus has the same position like the left-dorsal half of L4K of Tn-oniciis: 

 left-dorsal to the base of the hla-hook (compare fig. 85, 97 and 145. 151). In Tn'onicus 

 this sclerotisation has been regarded as an anterior part of the L41-region (fig.325g,h). 

 Sclerite L4P of Ciyptoc evens (fig. 151) probably coiTCsponds to that part of L4K of 

 Tryoniciis immediately anterior to the hla-base. The right-ventral half of L4K of 

 Tryonicus (fig.325g) with the anteriormost L41-region (the anterior extension to the 

 right) and the L4n-region (nla-sclerotisation) has been, like the nla-process itself lost 

 in Cn'ptocercus. 



Secondly, the same relations result from a comparison of the muscle insertions of 

 Cryptocercus and other species: 



- The 12 and 14 of Ciyptocercus (fig. 155, 156), Mantoida (fig.49, 50), and Eiuycotis 

 (fig. 70. 71) run from the pouches pne and Ive to the leftmost part of the left complex, 

 where their insertions are close to each other. Homology can be assumed. The left 

 insertions are on the L4l-region in Eunrotis and Mantoida (fig.325d,e), and on L4K 

 in Ciyptocercus. The contribution of the L41-region to L4K of Ciyptocercus is thus 

 confirmed (fig.325h). This can be only an anterior part of L41 since the posterior part 

 (with the pda-sclerotisation) is included in sclerite L4N. In Anaplecta L4K also bears 

 the left 12-insertion (fig.221). and in Nahiiblattella the homologue of the posterior part 

 of L4K (L4U') bears the left 12- and 14-insertions (fig.249). 



- The 114 of Ciyptocercus (fig. 157), Euiycotis (fig. 72), dind. Anaplecta (fig.222) run from 

 the anterior left wall of the left complex to a large hook (hla) and are certainly 

 homologous (discussion in 6.4.). Ciyptocercus (fig. 157) and Euiycotis (fig. 70) have 

 phallomero-sternal muscles sl+3 or si inserting immediately anterior to 114: the left 

 part of sl+3 {= si) is probably homologous with si of Euiycotis (si is missing in 

 Anaplecta: discussion in 6.9.). 



- In Euiycotis and Anaplecta 114 inserts on the L4n-region (on the nla-process, fig.72, 

 73, 222), and in Euiycotis si inserts at the border between L4n and the anterior L41 

 (fig.73, 325e). In Ciyptocercus part of 114 inserts on L4K; this suggests that the L4n- 

 region also contributes to L4K. The lai-ger part of the 114-insertion and the entire sl+3- 

 insertion, however, are on membrane (ventral and anterior to L4K); this suggests that 

 the L4ii-region as well as the anteroventral part of the L41-region (corresponding to the 



