229 



sclerotisation (fig. 222). (Homology is quite certain for s3 but not for 16a; discussion in 

 6.9. and 6.2.4.). That the dorsal insertion of s3 is on a sclerotisation is not comparable 

 with the situation in Lamproblatta and Polyphaga: In the latter species the sclerotisation 

 concerned is an expansion of the ventral plate (L4a-region); in Nahublattella the 

 sclerotisation with the s3-insertion is an expansion of the former L4K-sclerite. (The 

 respective area of the sclerite could be defined as a new sclerite region, but this is omitted). 

 Thus, L4U' is assumed to consist of the same parts of the L41-region as the posterior part 

 of L4K in Anaplecta. L4V' roughly corresponds to the L4n-region (fig. 325m, compare 

 fig. 3251); however, the line dividing the two sclerites does certainly not exactly correspond 

 to the border between L41 and L4n; this is only the case - by definition - in Archiblatta. 

 The homology relations of the processes paa and pda of Anaplecta and via, paa, pda, 

 and vsa of Nahublattella (fig.241, 244) and of their sclerotisations have been discussed 

 in 6.2.4.. L4N' of Nahublattella is probably the left- ventral sclerotisation of the via- 

 process (including pda and vsa; fig. 325m). The ribbon-like extension L4d' at the left base 

 of via (fig. 244, 250) closely resembles L4d of Polyphaga and Tryonicus (fig. 94, 97, 118, 

 129) in its position relative to paa and pda and their sclerotisation and to the UO-insertion. 

 Like in Polyphaga and Cryptocercus, L4d' is directed to the left. Like in Mantoida and 

 Cryptocercus, L4d' has a muscle running to the pne-pouch (11 in fig.48, 155, 249). 

 Homology is assumed for the L4d and 11 of all species. In Nahublattella the whole area 

 containing via, pda, paa, and L4d' is sunken anteriad into the left complex and has 

 become the left part of an expanded Ive-pouch, and L4d' Ues in the left edge of this 

 enlarged Ive-pouch and runs posteriad (L4d' is, so to speak, invaginated). This is in 

 contrast to all other species; only Anaplecta shows a slight anteriad invagination of the 

 paa+pda-sclerotisation (but L4d has been lost). The separation of the paa+pda- 

 sclerotisation from the L2-sclerotisations in the Ive-pouch reminds of Lamproblatta 

 (fig. 177- 179), but the division of the sclerotisations is different and non-homologous: L4d 

 is connected with the paa+pda-sclerotisation in Nahublattella but with the sclerotisation 

 of the Ive-pouch in Lamproblatta (compare fig.329f and g). 



The identification of the vla-lobe (fig.245-247) was done in in 6.2.4.. That there is no 

 sclerite plate in its ventral wall (L4v-region; compare L4G of Anaplecta, fig. 224) is a 

 derived feature. 



Parcoblatta, Blaberus, and other Blattelidae and Blaberidae 



Sclerite L4U' of Blaberus has the same shape, relative position, and muscle insertions as 

 L4U' of Nahublattella (fig.242, 249, 299, 303). In both species 14 runs to sclerite L2, and 

 12 runs to the hla-base (30 in fig. 249, 303). In Parcoblatta, the morphology of the left 

 part of the left complex (compare fig. 268-270 and 299-301) and the arrangement of 12 

 and 14 (compare fig. 276 and 303) are nearly the same as in Blaberus; however, sclerite 

 L4U has been lost. In Nyctibora L4U is present and very similar to L4U' of Blaberus. 

 Blaberus and Parcoblatta both have a tendon-like invagination (ate in fig. 268, 271, 302) 

 near the ventral basal line of the left complex, ate is also present, and in the same position, 

 in other Blattellidae and Blaberidae (investigated species: those listed in 5.15.; Blaptica: 

 fig.291). The homology of these ate-tendons is confirmed by the insertion of a phallomero- 



