234 



hla has a groove hge with a notch 45 in its ventral wall (fig. 266, 297a). In the species 

 studied only in part (listed in 5.15.), the hge-groove and the notch 45 are distinctly present 

 in Supella, EuphyUodromia, Loboptera, Byrsotria, and Blaptica; Nyctibora has only hge 

 but no notch 45; in Ectobius and Nauphoeta the hge-groove is quite indistinct, and the 

 notch 45 is missing. (114 and 136 have not been investigated in these species). Muscle 146 

 is pecuUar to Blaberus (fig.304, left part). 



6.5. Left complex V: Further main sclerites and muscles 



Some Blattaria and Mantodea have small sclerites in the dorsal wall of the vla-lobe, which 

 I have designated L5. L5 of Metallyticus (fig. 26, 27) and Cryptocercus (fig. 151, 155) is 

 posterior to the genital opening. L5 of Periplaneta (no figure) hes more anteriorly, within 

 the terminal part of the ejaculatory duct. L5 of Polyphaga (fig. 123, 124) is far to the left 

 of the genital opening. L5 of Ergaula is situated like in Polyphaga but is tranversely 

 orientated and approaches the genital opening more closely (fig.322m). In Anaplecta and 

 Nahublattella, the extension 28 of the L2- or L2D'-sclerite (fig.214, 215, 245) takes a 

 very similar position relative to the other parts of L2 and to the genital opening as L5 of 

 Polyphaga (fig. 123) and might be homologous with it. The sclerites LIO' of Blaberus and 

 Blaptica (fig.291, 300) lie either in the dorsal via- wall {Blaptica) or along the posterior 

 edge of the vla-lobe {Blaberus); whether they show any kind of homology relation with 

 the L5 of the other species is unclear, and improbable in my view. Sclerites in the dorsal 

 vla-wall are missing in Mantoida, Chaeteessa, Sphodromantis, Archiblatta, Eurycotis, Try- 

 onicus, Lamproblatta, and Parcoblatta. It cannot be decided whether L5 is a ground-plan 

 element of Blattaria and Mantodea and has been lost several times, or whether such 

 sclerites have developed several times independently. 



Sclerite L7 is present only in Polyphaga, Ergaula, and Lamproblatta. These L7 (fig. 115, 

 174) take the same relative position between the sclerite plate of the vla-lobe (L4M, L4R) 

 and the right phallomere and are therefore assumed to be homologous. L7 is regarded as 

 an element of the left complex since in a specimen of Polyphaga with its external genitalia 

 consisting of two right phallomeres only there was no trace of L7 (compare in 3.1.). Only 

 in Polyphaga and Ergaula the area containing L7 is elaborated as a special lobe-like 

 evagination (Iba in fig. 115; in Ergaula the morphology is the same, but L7 and Iba are 

 larger). The Iba-lobe is assumedly homologous with the rightmost part of the vla-lobe of 

 the other species. (If this is true, not the vla-lobe of Polyphaga and Ergaula alone but the 

 via- and Iba-lobes together are the strict homologue of the vla-lobe of the other species. 

 That Iba is not alone the homologue of the via of the other species and that L7 is not the 

 homologue of the L4G-plates is clearly shown by the muscles 15, 16a, and 16b, compare 

 in 6.2.1.. L7 and Iba are bare of muscles). 



Sclerite L8 is likewise restricted to Polyphaga, Ergaula, and Lamproblatta (L8 = 

 neoformation N of Grandcolas & Deleporte 1992). These L8 take the same position in 

 the right dorsal wall of the left complex, but they differ somewhat in their position relative 

 to the pne-pouch (fig. 117, in Ergaula similar; fig. 177). However, it must be considered 

 that in Lamproblatta, as compared with Polyphaga, the pne-pouch has shifted right- 



