246 



The age-apodeme is rather variable in its extension: In Polyphaga (fig. 137), Ergaula, 

 Anaplecta (fig. 229), Archiblatta, and Eurycotis (fig. 77), it is restricted to the right and the 

 right anterior margins of R3. In Cryptocercus (fig. 163) and Nahublattella (fig. 257) it 

 extends along the whole anterior and lateral margins of R3. In Parcoblatta (fig. 284), 

 Nyctibora, Byrsotria, and Blaberus (fig. 3 1 2a) it is restricted to the anterior marginal areas 

 of R3. Only in Tryonicus (fig. 102) and Lampwblatta (fig. 193) age has been lost. 

 R2 is an isolated sclerite in Eurycotis (fig. 75, 76), Tryonicus (fig. 100, 101), Cryptocercus 

 (fig.161, 162), Eamproblatta (fig.190, 191), Anaplecta (fig.227, 228), Nahublattella 

 (fig.254, 255), and Supella. In all these species, the right-ventral end of R2 articulates 

 distinctly with R3 (A7), the left-dorsal end of R2 articulates distinctly with Rl- 

 sclerotisations in the dorsal wall of the cbe-invagination (A6), and R2 is restricted to the 

 ventral base or, at least, to the ventral wall of the cbe-invagination. 

 In Euphyllodromia (fig.330o), Parcoblatta (fig.282, 283, 330q), Nyctibora (fig.330r), 

 Blaberus (fig. 3 10, 311, 330s), and Byrsotria R2 is fused to Rl-sclerotisations (RIS or 

 RIT) in the area corresponding to the A6-articulation of the other species. The point of 

 fusion is A6*, with the cwe-thickening (fig.282, 310) in its immediate vicinity. This topic 

 will be discussed in 6.7.6.. In Parcoblatta and Blaberus R2 is in close vicinity to R3 in 

 the area corresponding to the A7-articulation of the other species (A7 in fig. 284, 312a) 

 but is not articulated with R3. In Euphyllodromia and Byrsotria (fig.318) A7 is a distinct 

 articulation. In Nyctibora A7 is distinct and hinge-like (fig. 3 19). 



In Polyphaga (fig. 135- 137, 3301) and Ergaula (fig.330m) R2 and R3 are clearly 

 identifiable by their shapes (R3 is a broad curved plate, R2 forms a dental ridge) and by 

 their positions relative to each other, to cbe, and to the r2-insertions (fig. 141; compare 

 Cryptocercus, fig.161, 167, and Eurycotis, fig. 75, 81; r2 not investigated in Ergaula). 

 However, in both species R2 shows two peculiarities: (1) R2 is fused to R3. (2) R2 has 

 spread over the cbe-invagination (sclerotisation X in fig.3301,m) and is broadly fused to 

 Rl-sclerotisations in the dorsal cbe-wall; hence, the whole of cbe is sclerotised (fig. 134, 

 135). (1): In Polyphaga the stripe of weaker sclerodsation A7* (fig. 135, 137) takes the 

 same position as articulation A7 in other Blattaria and is assumed to be the line of fusion 

 between R2 and R3 (and a vestige of A7). In Ergaula R2 and R3 are fused without any 

 vestige of A7 (no weak line), and the border is not exactly determinable. Moreover, R2 

 of Ergaula has become so broad that R3 is for most of its breadth confluent with R2 

 (compare fig. 3301 and m). (2): In Polyphaga the sclerotisation of cbe has a weak line (13 

 in fig. 134, 138) and an adjacent notch within the sclerite margin; these structures are 

 assumed to mark the border between R2 and Rl, and the sclerodsation in the ventral wall 

 and on the summit of cbe is assumedly part of R2 (X in fig. 3301). Rl is restricted to the 

 posterior dorsal wall of cbe. In Ergaula Rl and R2 are firmly connected (no weak line); 

 the interpretation of the cbe-sclerotisation is done in accordance with Polyphaga (X in 

 fig.330m is part of R2). 



Archiblatta also has the whole cbe-invagination sclerotised (compare fig. 74 and 75 of 

 Eurycotis; the sclerotisation concerned is Y in fig.330f, compare fig.330g), but, in contrast 

 to Polyphaga, the cbe-sclerotisation is slightly weaker near the ventral base of the cbe- 

 invagination; in the corresponding area of Eurycotis R2 has its dorsal margin and the 



