250 



Eurycotis and Archiblatta 



For Eurycotis this area has been fully discussed in 6.7.1.. In Archiblatta, like in Eurycotis 

 and in all Mantodea, the fda-lobe and the pia-tooth are both very distinct. Sclerite RIF 

 (fig.330f, regions Rlc and Rlt) closely resembles RIF of Eurycotis. The sclerotisations 

 of the Rid- and Rlv-regions are more complicated than in Eurycotis (compare fig.330f 

 and g) but similarly structured in a dorsal (RIH = Rid) and a ventral (RIG = Rlv) 

 sclerite. The sclerite bridge connecting RIH and RIG in Eurycotis (behind membrane 17 

 in fig.77; fig.330g) has a short gap in Archiblatta; instead, there is a ribbon-like connection 

 between RIH and RIG across the ventral wall of the fda-lobe (compare fig.330f and g). 



Tryonicus, Cryptocercus, and Lamproblatta 



In these species (fig.99-104, 160-164, 190-195) the posterodorsal part of the right 

 phallomere has only two sclerites RIF and RIJ. RIF corresponds to RIF of Eurycotis 

 (fig. 74-78). The area posterior to RIF contains RIJ, which is a fusion product of RIH 

 and RIG of Eurycotis, and is an undivided lobe (fda, no ventral tooth pia present). The 

 regioning of Rl is shown in fig.331e,f,g,h and 332e,f,g,h. 



RIF (fig. 102, 163, 193) is in these species, like in Eurycotis, somewhat horseshoe-shaped 

 (open to the left, with a dorsal and a ventral arm), and along RIF there are the following 

 structures in common, which are all regarded as homologous: 



- The ventromedian end of RIF articulates with R2 (A6 in fig.75, 100, 160, 164, 190). 



- The ventral arm lies in the dorsal wall of the cbe-invagination (fig. 74, 99, 160, 190). 

 It bears a ridge (pva in fig. 80, 99, 164, 190, 196), which is formed by cuticular 

 evagination in Eurycotis, Tryonicus, and Lamproblatta, and by cuticular thickening in 

 Cryptocercus (cross-section in fig. 164). 



- At the base of this ventral arm the posterior margin of RIF articulates with sclerite RIJ 

 (A9 in fig. 102, 103, 190, 193), or the sclerites are at least in close vicinity (A9 in 

 fig. 163, 166). This corresponds to the position of articulation A9 of Eurycotis (compare 

 fig.77 and 78). Special features of Lamproblatta are the extension 20 of that part of 

 RIF bearing A9 and the immersion of the whole articulation. 



- From A9 RIF extends to articulation A3; then it curves into the dorsal wall of the right 

 phallomere. 



- At its dorsomedian end RIF has another articulation with RIJ (A8 in fig.99, 190) or, 

 at least, closely approaches RIJ (A8 in fig. 160). This corresponds to the position of 

 articulation A8 of Euncotis (fig. 74). 



- This dorsal arm of RIF has, like in Eurycotis, a sclerotised groove at its dorsal margin, 

 between the articulations A3 and AS (rge in fig.74, 77, 99, 102, 160, 163, 190, 193). 



RIJ (fig.99, 102-104, 160-164, 166, 190, 192-195, 197) bears both the articulations AS 

 (like RIH or region Rid in Eurycotis) and A9 (like RIG or region Rlv in Eurycotis) 

 and is therefore regarded as a compound sclerite Rld+Rlv (fig.331f,g,h, 332f,g,h). Thus, 

 in contrast to the situation in Mantodea, Eurycotis, and Archiblatta, there is now a very 

 broad connection between the Rid- and Rlv-regions posterior to membrane 17, and this 

 is clearly a derived feature. For Cryptocercus, Lamproblatta, and Eurycotis these relations 



