251 



are confiiTned by a comparison of the muscles, since RIJ bears insertions which are in 

 Eiuycotis either on RIH or on RIG: 



- Muscle r3 of Ciyptocercus and Lamproblatta (fig. 166, 197) runs from that part of RIF 

 posterior to articulation A3 to the right margin of RIJ. It is assumed to be homologous 

 with r3 of Eiuycotis (fig. 80), which inserts on RIG. 



- Muscle r6 of Lamproblatta (fig. 196) runs from the rge-groove to the dorsal wall of the 

 right phallomere, like r6 of Eurycotis (fig. 79). The left insertion is partly on RIJ in 

 Lamproblatta and on RIH in Eurycotis. Such a muscle is missing in Cryptocercus. 



- Muscle rl of Cryptocercus (fig. 165) runs from the age-apodeme on R3 to the dorsal 

 wall of the right phallomere, like rl of Eurycotis (fig.79). The posterior insertion is 

 partly on RIJ in Cryptocercus and on RIH in Eurycotis. Such a muscle is missing in 

 Latnproblatta. 



The fda-lobe of Tryonicus (fig.99, 102-104), Cryptocercus (fig. 160-164, 166), and 

 Lamproblatta (fig. 190- 195, 197) largely corresponds to fda of Eurycotis. However, parts 

 of its ventral wall assumedly correspond to the pia-walls of Eurycotis (after having been 

 leveled). Thus, the fda-lobes of these species are not strictly homologous with fda of 

 Eurycotis. The levelling of pia is also a derived feature. 



At least Lamproblatta has a similar flap-mechanism as Eurycotis (with fda being the flap 

 and the stout A8 and A9 defining the axis of movement). To what extent this is also 

 practised in Tryonicus and Cryptocercus is questionable since the articulations A8 and A9 

 are by far less distinct. 



Polyphaga and Ergaula 



In Polyphaga the posterodorsal part of the right phallomere contains the large sclerite 

 RIM and the smaller sclerites RIK and RIL (fig. 134). The regions Rlc, Rlt, Rid, and 

 Rlv can be identified and demarcated by their characteristic features (fig.331i, 332i), but 

 some points remain unclear. 



- RIM articulates with R3 (A3 in fig. 137) and forms a rge-groove on its dorsal margin 

 (from A3 to the posterior: fig. 134, 137, 140), and rge bears the insertion of a stout 

 muscle (r6 in fig. 140). These features resemble the Rlc-region of Eurycotis (fig.331e,i, 

 332e,i) and the other species. In contrast to the other species, the rge-groove extends 

 much farther posteriad (compare fig.74, 99, 160, 190). 



- To the left of A3 RIM bends around an edge (16 in fig. 137) to occupy the dorsal wall 

 of the cbe-invagination. The right part of muscle r2, coming from R3, inserts at the 

 anterior margin of this part of RIM (fig. 140). More to the left this part of RIM forms 

 a ridge (pva in fig. 139, 137, 138). These features resemble the Rlt-region of Eurycotis 

 (fig.331e,i, 332e,i) and the other species. In contrast to the other species, the pva-ridge 

 is not transversely but longitudinally orientated (compare pva in fig. 139 and 80, 99, 

 197). However, the shape of pva of Polyphaga is not so different from pva of 

 Lamproblatta (compare fig. 139 and 197), if a lengthening of pva along the longitudinal 

 axis and a shortening along the transverse axis is assumed for Polyphaga. 



- Corresponding to the probable lengthening of pva and rge to the posterior, it is assumed 



