254 



that in Polyphaga the regions Rlt and Rlc have considerably expanded posteriad and 

 make up most of RIM (fig.331i, 332i). 



- Muscle rl (fig. 139) has its anterior insertion on the right margin of R3, like rl and r5 

 of Eurycotis. The dorsal part of rl has its posterior insertion in the dorsal wall of the 

 right phallomere (on the RlL-sclerites), hke rl of Eurycotis (fig.79). This suggests that 

 the two RlL-sclerites are part of the Rld-region (fig.331i). The ventral part of rl has 

 its posterior insertion on the rge-groove (fig. 134, 139), similar to r5 of Eurycotis (fig.80) 

 but more posteriorly. Thus, rl of Polyphaga is certainly homologous with rl of 

 Eurycotis but possibly also includes the homologue of r5. rl of Cryptocercus closely 

 resembles rl of Polyphaga, but a contribution of a r5-part is less probable since no 

 fibers insert on rge or RIF (fig. 165). 



- Eurycotis (fig.79) and Lamproblatta (fig. 196) have the left insertion of r6 in the dorsal 

 wall of the fda-lobe and, by definition, in the Rld-region. r6 of Polyphaga has a very 

 similar course; its left insertion is on sclerite RIK, which is therefore assumed to belong 

 to the Rld-region (fig.331i). Thus, the Rld-sclerotisations of Polyphaga have become 

 rather fragmented (3 sclerites) and far removed from each other (as the insertions of rl 

 and r6 are). However, RIK and RIL could also be new elements not homologous with 

 sclerotisations of other Blattaria. RIK of Cryptocercus (fig. 160) could well be 

 homologous with RIK of Polyphaga. However, since r6 is missing in Cryptocercus, 

 the somewhat similar position of the sclerites is the only indication for homology. 



- Further parts of the regions Rid and Rlv might be included in the posterior part of 

 RIM. Compared with Cryptocercus (fig. 160, 163) or Lamproblatta (fig. 190, 193), this 

 would correspond to a fusion of RIF and RIJ across the membrane 17 and the 

 articulations A8 and A9. This is possibly indicated by the complete loss of muscle r3, 

 which in Cryptocercus (fig. 166) and Lamproblatta (fig. 196, 197) moves RIF and RIJ 

 upon each other: The loss of r3 could be the consequence of such a fusion. In fig. 3301, 

 33 li, and 332i Rid and Rlv are shown according to this assumption. 



The posterior part of the right phallomere is assumed to be composed of the fda-lobe and 

 of the pva-ridge (fig. 136- 138). Like in Lamproblatta, Cryptocercus, and Tryonicus, the 

 ventral tooth pia has been lost, and its leveled vestiges are assumed to be contained in 

 the ventral wall of fda. 



RIM of Ergaula is very similar to that of Polyphaga but narrower (compare fig. 3301 and 

 m). RIL and RIK, however, are missing. The regioning of RIM is assumed to be the 

 same as in Polyphaga. 



Anaplecta 



The posterodorsal part of the right phallomere is, like in Tryonicus, Cryptocercus, and 

 Lamproblatta, an undivided lobe (fda in fig. 226-230; no pia-tooth present). In contrast to 

 these species (with RIF and RIJ), however, there is only one sclerite present (RIN in 

 fig. 226-230), which somewhat resembles RIM of Polyphaga (fig. 134, 137). The regioning 

 of RIN is assumed to be as follows (fig. 33 Ik, 332k): 



- The part of RIN immediately posterior to articulation A3 (fig. 229) is the Rlc-region. 

 However, the rge-groove is missing. 



