255 



- The extension 34 of RIN, which to the left of A3 bends into the cbe-invagination 

 (compare fig. 229 and 230; fig. 226), exactly corresponds with the Rlt-region of the other 

 species by its relative position, by its articulation with R2 (A6 in fig. 226, 227, 230), 

 and by bearing the insertion of the right part of muscle r2 (fig. 231) (compare fig. 33 Ik 

 and 331f,h). Rlt of Anaplecta is somewhat thickened to the outside (pva) but does not 

 form a true ridge. 



- The posterior main part of RIN takes the same position as the RlJ-sclerite in Tryonicus, 

 Cryptocercus, and Lamproblatta and is probably composed of the regions Rid and Rlv. 



RIN is assumed to have developed by a fusion of the former RIF and RIJ across the 

 membrane 17 and the articulations A8 and A9 (compare fig. 160, 163 and 226, 229), as 

 it has also been assumed for RIM of Polyphaga (compare fig.331i and k, 332i and k). 

 Moreover, like in Polyphaga, muscle r3 has been lost. In contrast to Polyphaga, however, 

 the Rlt-region retains the same degree of independence and the same transverse 

 orientation as it has in e.g. Eurycotis and Tryonicus, and there are no free sclerites RIL 

 and RIK. Thus, it is not clear if RIN of Anaplecta and RIM of Polyphaga are strictly 

 homologous and if the fusion of the former RIF and RIJ and the loss of r3 are 

 homologous in the two species. (Therefore the sclerites are given different names). 

 The muscle connecting R3 and RIN (rl in fig.231) could be homologous with rl or r5 

 or both muscles of Eurycotis (fig.79, 80) and with the rl of Polyphaga (fig. 139) and the 

 other species. Like in Polyphaga, the muscle will be named rl in Anaplecta (and in the 

 other Blattellidae and Blaberidae discussed below). 



Nahublattella 



The posterodorsal part of the right phallomere is, like in Anaplecta, an undivided lobe 

 (fda in fig.253, 256: no ventral tooth pia) with one sclerite (RIN'), but the Rlt'- 

 sclerotisation seems to be missing (compare 34 in fig. 226). However, similar to the left 

 end of 34 in Anaplecta, the left end of RIN' (34 in fig. 253) articulates with R2' (A6 in 

 fig. 254, 255, 226) and curves back to the right like a hook. Therefore, the Rlt'-region is 

 assumed to have fused to the main part of RIN' lying posterior to it (fig. 33 11). Apart from 

 this difference, RIN' of Nahublattella is regioned in the same way as RIN of Anaplecta 

 (compare fig. 33 Ik and 1, 332k and 1). A peculiar feature of Nahublattella is the hinge-like 

 shape of articulation A3 (fig.253, 257). Muscle rl is certainly homologous with rl of 

 Anaplecta (fig.231, 259). Muscle rlO (fig. 259) is specific to Nahublattella. 



Supella 



The posterodorsal part of the right phallomere is again an undivided lobe fda with one 

 large sclerite RIN', but RIN' has expanded over the whole fda-lobe and over the whole 

 dorsal wall of the cbe-invagination. The Rlt'-region must have been firmly integrated into 

 this sclerotisation. Articulation A6, indicating the left end of the Rlt'-region, is distinct 

 and, like in Nahublattella, on the summit of che (compare fig.253, 254). A hook-like or 

 curved sclerotisation near A6, however, is not present. Supella resembles Nahublattella in 

 probably having Rlt' completely integrated into RIN', but because of the large expansion 

 of RIN' in Supella the situations in the two species are hardly comparable. 



