263 



The an-angement of the insertion areas on the subgenital plate (compare s4 in fig. 40 and 

 s4a and s4b in fig. 5) and on the phallomere complex (s4a and s4b insert medially like s4 

 in Mantoida, compare fig. 7 and 42) suggests that this is due to a division of s4. Both 

 species lack secondary muscles. 



Eurycotis 



si, s2, s3, and s4 conform with the basic arrangement (fig. 62, 64, 333d), but s2 is rather 

 weak. s5 and s6 have divided into three (s5a,b,c) or four (s6a,b,c,d) bundles, whose 

 insertions occupy a large area in the anterior half of the subgenital plate. However, most 

 of these bundles are very diffuse and indistinctly bordered to each other. Moreover, it is 

 not clear if really all these muscles are derivatives of s5 and s6 or if some of them are 

 new. Of the secondary muscles s7 and s8 (fig. 58) are present. 



Anaplecta 



The left side has three muscles (fig. 204, 207, 333h): The secondary s7 (fig. 200) runs, like 

 in Eurycotis, to the Ive-pouch. s5 can be identified by the lateral position of its ventral 

 insertion and by its dorsal insertion on the lateroventral wall of the genital-pouch. The 

 third muscle (named s3) might be si or s3. It has its ventral insertion posterior to s7 like 

 s3 of Eurycotis (si of Eurycotis inserts anterior to s7, compare fig. 333d and h) and has 

 its dorsal insertion immediately anterior to 16a (fig. 222) Hke s3 in Eurycotis (fig. 70, 73), 

 Polyphaga, and Lamproblatta (discussion below; fig. 133, 188). This muscle of Anaplecta 

 is therefore regarded as s3; si is missing. 



On the right side all three primary muscles are present (s2, s4, s6 in fig. 204, 207, 333h). 

 Their ventral insertions resemble the basic arrangement (s2 and s4 on the median anterior 

 margin; s2 anterior to s4; s6 far laterally), but the s4-insertion is very broad, and s2 is 

 strongly reduced. The insertions on R3 are side by side: s4 on the left, s2 in a small central 

 area, s6 on the right. Thus, in contrast to the basic arrangement, the dorsal insertion of s6 

 has expanded to the anterior right margin of R3 (but still occupies parts of the membranous 

 ventral wall of the genital-pouch as well). The same is true of s6a of Eurycotis (fig. 64, 

 333d), whose assignment to s6 is thus confirmed by the morphology of s6 of Anaplecta. 

 Noticeably, just si and s2 have been reduced - two muscles which probably are a pair. 

 Apart from s7 there is another secondary muscle: slO, which, however, is not directly 

 inserted on the ejaculatory duct but more ventrally on the infolding between the lobes via 

 and vfa (fig.221, 222; in fig.333h this infolding is symboUsed by a transverse line beneath 

 the ejaculatory duct; compare discussion in 6.2.4.). 



Lamproblatta and Polyphaga 



On the left side, si and s3 of Lamproblatta (fig. 173, 175, 333e) conform with the basic 

 arrangement, but the dorsal insertion of si is rather far on the left (compare Eurycotis, 

 fig. 70). In Polyphaga (fig. 11 3, 116, 333g) both insertions of s3 and the dorsal insertion 

 of si exactly correspond with Lamproblatta (si: membrane anterior to the hla-hook, 

 fig. 127, 185), but the ventral insertion of si has shifted leftward, too, and is not anterior 

 to but to the left of s3. Homology can certainly be assumed for si as well as for s3. The 



