271 



fused to the paraprocts laterally, would then have a more derived state of this character. 

 In the species having no separate Pv-sclerites by fusion to Pp (e.g. Mantoida, fig. 37, 

 Polyphaga, fig. 110) this character would be even more derived. The fusion of the 

 paraprocts Pp and the paratergites TlOp in Lamproblatta (partial; fig. 170), Anaplecta, 

 Mantoida, and Sphodromantis (complete; fig.201, 37, 2), corresponding to a partial 

 {Lamproblatta) or complete loss of A99, is certainly a derived feature. The presence of 

 two articulations per side is pecuhar to Lamproblatta (A97 and A99 in fig. 169, 170). The 

 interpretation in this species is done in accordance with Eiirycotis (fig. 59) and Tryonicus 

 (fig.83): The lateral articulation is the true A99; A97 is assumed to be a new articulation 

 within the paraproct Pp. 



In several species tergite 10 TIO has undergone a complete longitudinal division by a 

 median stripe of membrane {Polyphaga, fig. 109; Nahublattella, fig. 234; Blaberus, fig. 293) 

 - certainly a case of threefold parallel evolution. The membranous area 21 of Lamproblatta 

 (fig. 169) might represent an early stage of such a division. 



The articulation A98 between the cereal base and tergite 10 has been lost only in 

 Polyphaga and Blaberus - certainly another case of parallel evolution. 

 The ventral sclerotisation of tergite 10 TlOv is only in Anaplecta separated from the dorsal 

 main part of TIO (fig. 200). However, the TlOv-sclerites of Anaplecta could also be 

 homologous with the Cc-sclerites of the other species (compare fig. 200 and e.g. 58). 

 The various paired sclerites median to the cereal base (Ca, Cb, Cc) are certainly 

 homologous in the way expressed by the designations. All three pairs are present only in 

 (some) Blattaria but not in Mantoida and Sphodromantis {Chaeteessa and Metallyticus not 

 investigated). Sclerites median to the cereal base are also present in e.g. Caelifera 

 (Snodgrass 1935, fig. 7), but whether there is any kind of homology with the Blattarian 

 sclerites is unknown. It is therefore also unclear if some or all of these sclerites are 

 elements of the Dictyopteran ground-plan or derived features of Blattarian subgroups or 

 of Blattaria as a whole. Ca-sclerites are present in Eurycotis (fig. 58, 59), Tryonicus (fig.83, 

 84), Lamproblatta (fig. 169, 170), Anaplecta (fig.200), Nahublattella (fig.234, 235), and 

 Parcoblatta (fig. 262, 263), and they are crescent-shaped in most species. Except in 

 Tryonicus and Lamproblatta the Ca extend along distinct curved Ca-bulges. Cc-sclerites 

 are present in Eurycotis (fig.58, 59), Tryonicus (fig.83, 84), Lamproblatta (fig. 169, 170), 

 and possibly Anaplecta (fig.200: TlOv?). Cb-sclerites are peculiar to Lamproblatta 

 (fig. 169, 170). In Polyphaga, Cryptocercus, and Blaberus all three pairs are missing, but 

 in Polyphaga and Blaberus at least the Ca-bulges are distinct. 



A distinct supraanal lobe spl has been found in Mantodea (fig.l, 36) and in Eurycotis, 

 Tryonicus, Cryptocercus, Lamproblatta, Parcoblatta, and Blaberus (fig.58, 83, 143a, 169, 

 262, 293). An epiproct Ep is present in Mantodea (fig.l, 36) but never in Blattaria. 



6.13. The asymmetry of the phallomere complex 



The right phallomeres of the Mantodean species, especially Chaeteessa, and of Eurycotis 

 are very similar in the arrangement of the sclerotisations (Rl, R2), the formative elements 

 (invagination cbe, lobe fda, tooth pia, ridge pva, apodeme age), the main muscles (rl, 



