279 



Subgroup 1.1.: Mantoida 



The phallomere complex of Mantoida is rather close to the Mantodean ground-plan but 

 also has some derived features: On the left complex, the L4n-region has been lost (6.3.1., 

 6.3.3.). On the right phallomere, the Rlt-region (with the pva-tooth) has been separated 

 from the Rlc-region (6.7.1., 6.7.3.; like in Metallyticus: compare (G) in 7.5. and grouping 

 B (123) in 7.6.). 



Subgroup 1.2.: Chaeteessa + (Metallyticus + Sphodromantis) 



There are some conspicuous autapomorphies on the left complex: The ventral wall has 

 developed the L4b-region which occupies all the interspaces between the primary L4- 

 regions L4v/L4c, L4n, and L41 and makes the ventral wall completely, or nearly so, 

 sclerotised (6.3.3.; in the primitive case L4b is distinctly weaker than the primary L4- 

 regions). The swe-apodeme has been lost or reduced to vestiges (6.3.3.). In the dorsal wall 

 the L41-region has strongly expanded to the right (6.3.3.). The L4d-region, if present at 

 all, is no longer prominent from the outline of the L41-region (6.3.3.). Possibly in 

 correlation with this expansion of L4I, the pne-pouch has rotated (clockwise as seen from 

 behind), and the membranous part of its wall is somewhat more on its right side (6.1.3.). 

 The distal part of the right posterior extension Llm of sclerite LI curves into the dorsal 

 wall of the Ive-pouch, and Llm and L2 are therefore in the same plane in the area of 

 articulation A2 (6.1.3.). The Ive-pouch has become distinctly narrower (6.2.3.). Sclerite 

 L2 in its dorsal wall has lost its primitive arch-shape (probably by a fusion of the arms 

 of the arch, L2p and L2m) and is now ribbon- or plate-like (6.2.3.; compare (B) in 7.5.). 

 At least the paa-process has distinctly lengthened (6.2.3.). Whether the pda-process has 

 lengthened is not assessable since pda has been lost in Chaeteessa. The sclerotisations of 

 pda and paa (or, to apply this character state also to Chaeteessa, the L4- and L2- 

 sclerotisations of the corresponding area) have been separated from each other (6.2.3.; 

 compare (A) in 7.5.). 



On the right phallomere, the Rlc-region has been divided by the membranous stripe 4 

 separating the sclerites RIA and RIB (RIA and RIC in Metallyticus) (6.7.1., 6.7.3.). 



Subgroup 1.2.1.: Chaeteessa 



The left complex has a membranous pouch pbe between the pne- and Ive-pouches (fig. 34). 

 The pda-process has been lost (6.2.3., 6.3.3.). The loa-process has also been lost (assuming 

 that at least the loa of Mantoida and Metallyticus + Sphodromantis are homologous; 

 6.1.3.). The vla-lobe has considerably broadened as compared with the narrow Ive-pouch 

 (fig. 32), and the genital opening has come into a position far to the right of the Ive-pouch 

 (6.2.3.). On the right phallomere, sclerite R3 bends to the right along its left margin (6.7.3., 

 fig. 32). The keel-apodeme 3 has been lost (6.7.3.). The utmost right-posterior part of the 

 age-groove (near articulation A3) has been reduced (6.7.3.). 



Subgroup 1.2.2.: Metallyticus + Sphodromantis 



Most of the autapomorphies are on the left complex: The dorsal and ventral parts of L4 

 have been separated by articulation Al at the left edge of the left complex, which divides 



