285 



Subgroup 2.2.2.1.: Ciyptocercus 



The left complex has several derived features: The right part of L2, which is upcurved in 

 other Blattaria, has been reduced; in consequence, the respective right-dorsal part of the 

 Ive-pouch has been reduced, and the contact between L2 and LI (articulation A2) has 

 been lost (6.1.4., 6.2.4.). The left edge 61 of the vla-lobe has expanded almost to the 

 anterior margin of the left complex; in consequence, the left-ventral part of the Ive-pouch 

 has also been extremely reduced (6.2.4.; a similar derived state is present in Tryonicus 

 and in Anaplecta: compare grouping C (124) in 7.6.). The pda-sclerotisation (posterior 

 L41-region) has been reduced and also separated from the paa-sclerotisation (L2d-region) 

 (6.2.4., 6.3.4.; the separation of L41 and L2d has also developed in the subgroups 1.2. 

 and 2.1.: compare (A) in 7.5.). New muscles of Cryptocercus are 119 (6.4.3.), r7 (6.7.5.), 

 r8 (6.7.4.), and possibly 17 (6.5., 7.1.). 



There are some derived features in the phallomero-stemal and peripheral musculature: si 

 and s3, and probably s7, are fused (6.9.). s4 and s6, and s2 if present, are fused (6.9.). 

 The pi are extremely broad. The posterior insertions of the p4 are far laterally (6.11.). 

 All these seemingly primitive features are assumed to be neotenic traits; that they are not 

 primitive but derived results from the outgroup comparison with Mantodea. s5 has been 

 lost (6.9.; like in Polyphaga and Ergaula: compare grouping H (131) in 7.6.). 



Subgroup 2.2.2.2.: Lampwblatta + (Ergaula + Polyphaga) 



There are many autapomorphies on the left complex: L8 is a new sclerite in the right 

 dorsal wall, with the insertions of 112, 19, and b2 upon it or in its immediate vicinity (6.5.). 

 L7 is a new sclerite on the right part of the vla-lobe (or on the Iba-lobe, which is a right 

 part of via; 6.5.). The muscles 111 (6.3.4.), 112 (6.2.4., 6.5.), and sl2 (6.2.4., 6.3.4., 6.9.) 

 are also peculiar to this subgroup. The L4-plate in the ventral vla-wall has expanded to 

 include the dorsal insertion of muscle s3 (new region L4a, larger sclerites L4R and L4M; 

 6.3.4.). The left insertion of 12 has shifted further anteriad and away from sclerite L4K 

 (or region L41; 6.3.4.). The Ive-pouch has expanded almost to the left edge of the left 

 complex (6.2.1., 6.2.4.). 



Other derived features could possibly also be autapomoiphies of this subgroup: If s2-parts 

 should be included in s2+4+6 of Cryptocercus, s2 would have been lost in subgroup 

 2.2.2.2. (6.9.). The dorsal insertion of muscle s6 has become restricted to the anterior 

 margin of sclerite R3 (6.9.). (If the muscle named s6 should be s2, the loss of s6 and the 

 laterad expansion of the ventral s2-insertion would be the autapomorphies). If s7 really is 

 a ground-plan muscle of Blattaria (compare (L) in 7.5.), and if vestiges of s7 are included 

 in sl+3 of Cryptocercus, the complete loss of s7 would also be an autapomorphy of 

 subgroup 2.2.2.2. (6.9.). Muscle 11 has been lost. (However, 11 has certainly been lost 

 several times: at least also in the subgroups 2.1., 2.2.3.1., and 2.2.3.2.2.; compare grouping 

 R (25) in 7.6.) 



Subgroup 2.2.2.2.1.: Lamproblatta 



There are many derived features on the left complex. L2 has divided into three sclerites: 

 L2A in the left part of the Ive-pouch, L2B in the right part of the Ive-pouch, and L2C 



