288 



The following features have been investigated only in Anaplecta, Nahublattella, 

 Parcoblatta, and Blaberus: Muscle si has been lost (6.9.). The muscles termed 15 in 

 Anaplecta and Nahublattella are possibly new muscles, or they are true 15 with the anterior 

 insertion shifted to the L4n-region; in any case, one of these character states is probably 

 apomoiphic for subgroup 2.2.3. (6.2.4., 6.3.4.). (In Parcoblatta and Blaberus 15 has been 

 lost or integrated into 16b). 



Subgroup 2.2.3.1.: Anaplecta 



On the left complex, sclerite LI and the dca-processes have been lost (like in all or many 

 species of subgroup 2.2.3.2.2., compare grouping L (109) and (110) in 7.6.; according to 

 McKittrick (1964), however, LI is present in another species of Anaplecta; 6.1.4.). The 

 pne-pouch is therefore completely membranous; it has been reduced to a shallow 

 depression in the central dorsal wall, (pne has been lost completely in all or many species 

 of subgroup 2.2.3.2.2., compare (111) in 7.4. and grouping L (134) in 7.6.; 6.1.4.). Muscle 

 14 has been lost (6.2.4., 6.3.4.; like in Lamproblatta: compare grouping K (133) in 7.6.). 

 Some membranous foldings have developed in the area of the Ive-pouch, e.g. vfa and vpe 

 (6.2.4.). The gta-process (6.2.4., fig.215) and the vte-tendon (fig.208) have evolved. The 

 L4d-region has been lost (like in subgroup 2.2.3.2.2., compare grouping L (95) in 7.6.). 

 Muscle 11 has been lost (like in some other subgroups: compare groupings L and R (25) 

 in 7.6.). 125 and 126 are probably muscles peculiar to Anaplecta. (However, 126 might be 

 homologous with 16a of Nahublattella, Parcoblatta, and Blaberus; 6.3.4.). 

 The firm connection between the nla-process and the top of the Ive-apodeme might be 

 autapomorphic for Anaplecta. According to its possible con-elation with the translocation 

 of the anterior 114-insertion from nla to the top of Ive, however, this feature could also 

 be an autapomorphy of the whole subgroup 2.2.3., lost again at the base of subgroup 

 2.2.3.2. (compare (M), (N) in 7.5.; 6.4.3.). That there are two phallomere gland openings 

 in a rather peculiar position - possibly new organs - could also be either a derived feature 

 of Anaplecta alone or an autapomorphy of subgroup 2.2.3., with a loss of one opening 

 and a deplacement of the other at the base of subgroup 2.2.3.2. (6.6.). 



Subgroup 2.2.3.2.: Nahublattella + (Supella + (Euphyllodromia + (Parcoblatta + 

 (Nyctibora + (Blaberus + Nauphoeta + Blaptica + Byrsotria))))) 



At least the following apomorphies are present in all species listed: The anterior insertion 

 of muscle 114 has been translocated from the L4n-region on the nla-process to the L2a- 

 region on top of the Ive-apodeme (6.4.3.; or, if the homology of the 114-muscles should 

 not be true, a new muscle from L2a to L3 has developed). The size of the vla-lobe has 

 distinctly decreased (6.2.4., 6.3.4.). The common base of the paa- and pda-processes has 

 been elongated and forms, together with paa and pda, the via-process (6.2.4., 6.3.4.). The 

 right posterior dorsal part of the left complex - the part dorsal to the right half of the Ive- 

 pouch - has been reduced (6.2.4.). The division of L2 into L2D (within the Ive-pouch) 

 and L2E (together with L4N on the via-process) by articulation A 10 can also be regarded 

 as an autapomorphy of this group since this situation is present in both the subordinate 



