289 



sister groups 2.2.3.2.1. and 2.2.3.2.2. (The absence of this division and of AlO is regarded 

 as a secondary loss having occurred several times within subgroup 2.2.3.2.2.: compare 

 (Q) in 7.5.). 



The following derived features are also assumed to be autapomorphies of subgroup 2.2.3.2. 

 but have been investigated in sufficient detail only in Nahublattella, Parcoblatta, and 

 Blaberus: On the left complex, L4K has divided into the sclerites L4U and L4V - with 

 the nla-process probably still present on L4V' of Nahublattella. (In Blaberus and possibly 

 also in Parcoblatta L4V has been completely lost; 6.3.4.). Sclerite L4G (L4v-region) in 

 the ventral vla-wall has been lost (6.3.4.). The right insertion of muscle 12 has shifted to 

 the membranous basal part 30 of the hla-hook (6.3.4.). Muscle 130 has developed, having 

 a longitudinal course in the ventral wall of the left complex (6.5.). Muscle s2 has been 

 lost (6.9.; like in subgroup 2.2.2. or 2.2.2.2.: compare above). The muscles s5 and s6 have 

 divided into s5a and s5b or s6a and s6b, respectively (or, new muscles s5a and s6a have 

 developed; 6.9.). The p4-insertions on the subgenital plate have shifted mesad (6.11.). 



Subgroup 2.2.3.2.1.: Nahublattella 



On the left complex, the sclerotisation L2E'+L4N' of the via-process has been divided 

 transversely by the membrane ring 39 (6.2.4.). The nla-process, if actually nla, has become 

 whip-shaped (6.3.4.). The central dorsal wall contains a bristle area 35 (fig. 242). The 

 muscles 127, 128, 129, 131, 132, and 135 have developed (fig.240, 249-252). On the right 

 phallomere, sclerite R2 has become plate-like, bearing the highly elaborated evaginations 

 42 and 43 (6.7.4.). The Rlt-region has probably completely fused to the rest of sclerite 

 RIN' (6.7.6.). There is a new muscle rlO (6.7.6.). The dorsal sclerotisation S9d of the 

 subgenital plate is highly elaborated, divided, and provided with the new muscles 133 and 

 134 (6.10.). 



Subgroup 2.2.3.2.2.: Supella + (Euphyllodromia + (Parcoblatta + (Nyctibora + (Blaberus 

 + Nauphoeta + Blaptica + Byrsotria)))) 



At least the following apomorphies are present in all species listed (with the exceptions 

 mentioned): On the left complex, the hla-hook has evolved a hge-groove with a notch 45 

 in its ventral wall (6.4.3.). (In Nyctibora and Nauphoeta hge is not that distinct, and the 

 notch 45 is missing; this is probably due to secondary reduction). The ate-tendon has 

 become very long and narrow (6.3.4.). (That ate is rather short in Blaberus, Blaptica, and 

 Byrsotria is interpreted as a secondary reduction since ate is very long and thin in 

 Nauphoeta, and since these four species form the holophyletic subgroup 2.2.3.2.2.2.2.2.2.). 

 Sclerite L4V (essentially the L4n-region) has been reduced to a small sclerite in the dorsal 

 wall of the ate-tendon or has been completely lost (or, L4V has been generally lost, and 

 the sclerite within ate, present only in Parcoblatta, Nyctibora, and Blaptica, is a new one; 

 6.3.4.). The nla-process has been lost (6.3.4.). The via-process is no longer branched, i.e. 

 paa and pda are no longer distinct (6.2.4.). The right posterior branch of L2, whose distal 

 part sclerotises the psa-process in Nahublattella, has been completely lost (6.2.4.). The 

 L4d-region has been lost (6.3.4.; like in Anaplecta: compare grouping L (95) in 7.6.). The 



