290 



ventral extension 28 of L2, present in Anaplecta and Nahublattella, is lacking; this is also 

 assumed to be a derived feature (6.2.4.; compare the possible homology of 28 and L5: 

 6.5.). 



The following apomorphies have been investigated only in Parcoblatta and Blaberus; they 

 might be autapomorphies of the whole subgroup 2.2.3.2.2. or of any subordinate subgroup 

 containing at least Parcoblatta and Blaberus: The age-groove has been restricted to the 

 anterior part of sclerite R3, i.e. the age-part along the posterior right margin of R3 has 

 been lost (6.7.4.). Muscle 114 has divided into two bundles 114a and 114b (6.4.3.; this 

 division is completely different from the division of 114 in Eurycotis). Muscle 13 has been 

 lost (6.1.4., 6.2.4.). The muscles 136, 137, and 138 have developed (fig.276-278, 303, 307; 

 6.4.3.). Muscle 15 has been lost or integrated into 16b (6.2.4.). Muscle 16a has distinctly 

 enlarged (6.2.4.). Muscle s3 has divided into two bundles s3a and s3b (6.9.). The muscles 

 sl4 or sl4a,b are new (6.9.). The dorsal insertion of muscle s6b has expanded posteriad 

 to occupy the entire right margin of sclerite R3 (6.9.). The ventral insertions of s5a and 

 s6a have shifted posteriad (6.9.). Some further derived features of Parcoblatta and 

 Blaberus are also present in Anaplecta but not in Nahublattella (compare grouping L (109), 

 (110), (134), and (25) in 7.6.): Sclerite LI, the dca-process(es), and muscle 11 have been 

 lost. The pne-pouch, which has become indistinct in Anaplecta, has been completely lost 

 (6.1.4.). 



Subgroup 2.2.3.2.2.1.: Supella 



Sclerite RIN' has extremely expanded to occupy the entire dorsal wall of the cbe- 

 invagination (6.7.6.). The indistinctness of the Rlt'-region and the lack of a hook-hke 

 curvature at the median end of Rlt' (present in the ground-plan of subgroup 2.2.3.) are 

 assumed to be results of this expansion. 



Subgroup 2.2.3.2.2.2.: Euphyllodromia + (Parcoblatta + (Nyctibora + (Blaberus + 

 Nauphoeta + Blaptica + Byrsotria))) 



All species listed share some very important derived features on the right phallomere 

 (6.7.6.): The median ends of the Rlt-region and of sclerite R2 have fused: loss of 

 articulation A6. (A6 is still an articulation in Supella). The hook-curvature at the median 

 end of Rlt has been elaborated as the cwe-thickening. At its lateral end, Rlt has been 

 separated from the Rlc-region: resulting sclerites RIP and RIS. (In Supella Rlc and Rlt 

 are still connected with each other. In Blaberus, Nauphoeta, Blaptica, and Byrsotria Rlt 

 and Rlc are also connected: sclerite RIT'; this situation is interpreted as a secondary 

 fusion of these regions and as a synapomorphy of these species, which view is suggested 

 by the assumed autapomorphies of the subgroups 2.2.3.2.2.2.2. and 2.2.3.2.2.2.2.2.). 



Subgroup 2.2.3.2.2.2.1.: Euphyllodromia 



No derived features restricted to this species have so far been found in the phallomere 

 complex. According to the phylogenetic hypothesis presented here, the following derived 

 features appear as autapomorphies: The L2-sclerotisations of the via-process and of the 

 Ive-pouch have fused secondarily (i.e. articulation AlO has been lost). Muscle 110 has 



