291 



been lost. (Both features also in some other species of subgroup 2.2.3.2.2.2.; compare (Q), 

 (R) in 7.5.). The membranous left wall of the Ive-pouch has deeply invaginated to the left 

 (also in Loboptera; 6.2.4.). 



Subgroup 2.2.3.2.2.2.2.: Parcoblatta + (Nyctibora + (Blaberus + Nauphoeta + Blaptica 

 + Byrsotria)) 



Only two possible autapomorphies are present in the phallomere complex: First, the 

 presence of a dla-lobe (6.7.6.). dla is clearly missing in Supella and Euphyllodromia. dia 

 is also missing in Nauphoeta, but in this species the whole right phallomere has been 

 strongly reduced. Second, the presence of a tve-tendon (6.2.4.). tve is absent in Supella 

 and Euphyllodromia and present in all members of this subgroup except Blaberus 

 {Byrsotria not investigated). The absence in Blaberus is regarded as a secondary loss. The 

 holophyly of this subgroup is strongly supported by a clearly derived feature of the 

 females: They perform a rotation of the ootheca within the vestibulum (into a horizontal 

 orientation; McKittrick 1964; termed advanced rotation by Roth 1967). In Supella and 

 Euphyllodromia the ootheca retains a vertical orientation till it is dropped (Roth 1967). 



Subgroup 2.2.3.2.2.2.2.1.: Parcoblatta 



Features of Parcoblatta which are derived as compared with all other investigated species 

 are the vge-groove on the via-process (fig. 273) and the rotation of the via-process and of 

 some adjacent elements, which includes a ventrad shift of the genital opening (6.2.4.). 

 Some other features of Parcoblatta are derived at least as compared with Blaberus 

 (characters mostly not investigated in the other species of subgroup 2.2.3.2.2.): Sclerite 

 L4U has been lost (6.3.4.). Sclerite R2 has become distinctly curved (6.7.4.). The muscles 

 139 and 140 have developed (fig.277, 278). Muscle sl4 has divided into two bundles sl4a 

 and sl4b (6.9.). The ventral insertion of muscle 16a and the dorsal and ventral insertions 

 of muscles s5a and s6a have shifted very far posteriad (6.2.4., 6.9.). Further derived 

 features are the fusion of the L2-sclerotisations of the via-process and of the Ive-pouch 

 and the loss of muscle 110 (6.2.4.; both like in Euphyllodromia and Blaberus: compare 

 (Q), (R) in 7.5.). 



Subgroup 2.2.3.2.2.2.2.2.: Nyctibora + {Blaberus + Nauphoeta + Blaptica + Byrsotria) 

 This subgroup has some probably derived features on the right phallomere: There are two 

 new sclerites, R5 in the ventral part of the right phallomere (6.7.4.) and R4 in the dorsal 

 wall of the dla-lobe (6.7.6.). A new muscle rll runs from the dorsal dla-wall to the ventral 

 fda-wall (6.7.6.). 



Subgroup 2.2.3.2.2.2.2.2.1.: Nyctibora 



No certain autapomorphies have so far been found in the phallomere complex. 



Subgroup 2.2.3.2.2.2.2.2.2.: Blaberus + Nauphoeta + Blaptica + Byrsotria 



In all listed members of this subgroup, which corresponds to the Blaberidae sensu 



McKittrick (1964), the Rlt'-region (sclerite RIS') and the rest of Rl' (sclerite RIP') have 



