310 



Grouping E: Lamproblatta + (Anaplecta + (Nahublattella + (Supella + (Euphyllodromia 

 + (Parcoblatta + (Nyctibora + (Blaberus + Nauphoeta + Blaptica + Byrsotria))))))) 

 (73) Tendon tre and muscles s8 and b4a,b absent (SG2.: tre, s8, and b4a,b present; BM: 

 Presence of tre and s8 unclear, b4a,b present). 



This is probably the apomorphic state (compare (I) in 7.5.; 6.7.5., 6.9.). To regard it as 

 an autapomorphy of this grouping would be inconsistent with the many assumed 

 autapomorphies of the subgroups 2.2.2. and 2.2.2.2. 



Grouping F: {Polyphaga + Ergaula) + (Anaplecta + (Nahublattella + (Supella + 

 (Euphyllodromia + (Parcoblatta + (Nyctibora + (Blaberus + Nauphoeta + Blaptica + 

 Byrsotria))))))) 



(128) Region Rlc probably fused with at least part of the broadly interconnected regions 

 Rid and Rlv: articulations AS and A9 and membranous area 17 absent; sclerites RIM or 

 RIN (SG2.2.: Rlc separated from the broadly interconnected Rid and Rlv: AS, A9, and 

 17 present; SG.2.: Rlc separated from the narrowly interconnected Rid and Rlv: AS, A9, 

 and 17 present; BM: Rlc connected with both Rid and Rlv which are separated from 

 each other: AS and A9 absent but 17 present) (64) Muscle r3 absent: from region Rlc to 

 region Rlv (BM: r3 present). 



(128) is a less specific formulation of (63) as well as (74) (compare list in 7.4.) and is 

 true of both the subgroups 2.2.2.2.2. and 2.2.3. (128) and (64) are certainly apomorphic 

 - and the characters are probably intercorrelated (6.7.6.; compare (H) in 7.5.). The special 

 kind of fusion, however, is rather different in Polyphaga + Ergaula (subgroup 2.2.2.2.2.) 

 and in the other species (subgroup 2.2.3.); consequently, it is highly questionable whether 

 the fusion, the resulting sclerites RIM and RIN, and the loss of r3 are each homologous. 

 If these states are still regarded as autapomorphies of this grouping, these characters would 

 be inconsistent with the many assumed autapomorphies of the subgroups 2.2.2. and 2.2.2.2. 



Grouping G: Tryonicus + Lamproblatta 



(129) Apodeme age absent (BM: age present) (130) Extension R2m present (SG2.: R2m 

 absent). 



Both states are certainly apomorphic (6.7.4.). To regard them as autapomorphies of this 

 grouping would be inconsistent with the many assumed autapomorphies of the subgroups 

 2.2.2. and 2.2.2.2. 



Grouping H: Cryptocercus + (Polyphaga + Ergaula) 



(131) Muscle s5 absent: from subgenital plate to left wall of genital pouch (BM: s5 

 present). 



This state is certainly apomorphic (6.9.). To regard it as an autapomorphy of this grouping 

 would be inconsistent with the assumed autapomorphies of subgroup 2.2.2.2. 



Grouping J: Cryptocercus + Polyphaga 



(132) Sclerite RIK present (BM: RIK absent). 



The presence of a separate RIK is certainly apomorphic. However, the homology of the 



