334 



species (compare in 7.3., subgroup 2.2.2.; secondary reduction has been assumed for 

 iMmproblatta). 



- "Sclerite LI* turned on the ejaculatory duct and its opening." 



- "Apical apodema of sclerite LI* curved around the ejaculatory duct opening." 

 These two features probably refer to the hood-hke shape of sclerite LI and to its close 

 relation to the opening of the phallomere-gland (= "ejaculatory duct": compare in (A)). 

 Both features, however, are present in the common ground-plan of Blattaria and 

 Mantodea (6.1.1., 7.1.). 



- "Sclerite R2* with two tubercles, which are not articulated dorso-ventrally." 

 Misidentification (C). 



- "Female paraprocts with a membranous area in their sub-basal and internal parts." 

 According to fig. 13 of Grandcolas (= fig. 334a in this paper) this refers to a membranous 

 area at the median base of the subanal lobes (similar to Y in fig.321c). However, a 

 membranous area taking the same position is also present in the females of e.g. 

 Periplaneta (fig.334e), Deropeltis (fig.334d), and Lamproblatta (fig. 334c) and is 

 certainly not an autapomorphy of Polyphagidae or Polyphagidae -i- Cryptocercus. 



- "Straight, long and naiTow paratergites." According to my own investigations (Klass, 

 in press: te and tg in fig. 15, 16), the fused paratergites of the abdominal segments 8 

 and 9 of the females are, as compared with Periplaneta, somewhat lengthened and 

 narrowed in Cryptocercus but not in Polyphaga. Apart from this, a slight change of the 

 proportions of sclerite elements is in my view not very convincing as an autapomorphy. 



- "LI* pourvu d'une dilatation basale" (Grandcolas & Deleporte 1992). This feature 

 probably relates to the transverse expansion of LI at its posterior margin, which 

 continues towards both sides into the extensions Lll and Llm (fig. 120, 153, 323i,l). 

 However, a similar expansion, with at least one extension Llm, is also present in e.g. 

 Mantoida (fig.49, 323d), and this is certainly a feature of the common ground-plan of 

 Blattaria and Mantodea (6.1.1.). The extensions or regions Lll and Llr are also not 

 restricted to Cryptocercus and Polyphagidae (compare e.g. Tryonicus angustus, fig. 107, 

 323h, and Nahublattella, fig.243, 244, 323n). 



a) Polyphaga b) Cryptocercus c) Lamproblatta d) Deropeltis e) Periplaneta 



aegyptiaca pimctulatus albipalpus sp. americana 



334 



Fig. 334: Paraprocts of female Blattaria. - Ventral view of left subanal lobe; posterior?, anterior^, 

 median-^. Sclerotised areas (paraproct) are stippled, membranous areas are white. Fig. 334a according 

 to Grandcolas (1994). 



