132 



Fabian Herder, Jan Huylebrouck & Klaus Busse 



Type material in other collections: Holotype: IBTS 4587, 

 same data as ZFMK 22 1 5 1-22 1 52; Paratypes: IBTS 4586 

 (30), IBTS 4588 (6), ICDD uncat. (54), MGAB uncat. 

 (51). 



Remarks: ZFMK 22151-22152 are out of IBTS 4588. 



OSPHRONEMIDAE 



Betta climacura Vierke, 1988 

 Das Aquarium 22: 336 



Holotype: ZFMK 15532, male, 66 mm, small clearwater 

 brooks about 60 air-miles southwest of Brunei, near Ram- 

 payoh, West Borneo, Jul 1983, T. Schulz. 



Paratype: ZFMK 15533 (1), female, 65 mm, same data as 

 holotype. 



Current status: Synonym of Betta akarensis Regan, 1910 

 (Kottelat et al. 1993: 161). 



Betta persephone Schaller, 1986 

 AquarTerrarZ 39: 298 



Holotype: ZFMK 14226, male, Asian highway no. 2, 3 km 

 north of Ayer Hitam, Malaysia, 20 Feb 1985, D. Schaller. 



Paratypes: ZFMK 14227-14238 (12), same data as holo- 

 type. 



Type material in other collections: ZMB 31614 (4); 

 ZSM/CMK 5944 (4); same data as holotype. 



Betta tussyae Schaller, 1985 

 Aquar Terrar Z 38: 350 



Paratypes: ZFMK 14224-14225 (2), about 77 km south 

 of Kuantan at the road parallel to east coast, 1 7 km south 

 of Pekan, about 1 km from sea, Pahang State, Malaysia, 

 1984, P. Nagy; ZFMK 14195 (1), male, 16 km south of 

 Kuantan, Malaysia, 22 Feb 1985, D. Schaller; ZFMK 

 14196-14207 (12), same data as ZFMK 14195; ZFMK 

 14208-14212 (5), same data as ZFMK 14195. 



Type material in other collections: Holotype: ZRC 38451 

 (ex ZSM 27336); Paratypes: ZSM/CMK 5945 (5); 

 ZSM/CMK 6345 (11); ZMB 31607 (6); ZMB 31609 (1). 



Remarks: Originally the specimen ZFMK 14195 was de- 

 posited as holotype by the author in Bonn, but it cannot 

 be identical to the specimen which was characterized as 



Bonn zoological Bulletin 59: 109-136 



holotype by measurements in the original description be- 

 cause the ZFMK specimen is smaller. Nevertheless this 

 specimen belongs to the series of paratypes. 



Macropodus erythropterus Freyhof & Herder, 2002 

 Ichthyol Explor Freshwaters 13:162 



Holotype: ZFMK 38945, 58.9 mm, stream at Quyet Tien, 

 Quang Binh Prov., Vietnam, 17°11.54'N 106°37.53'E, 8 

 Apr 2000, J. Freyhof & F. Herder. 



Paratypes: ZFMK 38946-38961 (16), 23.7-57.3, same da- 

 ta as holotype; ZFMK 38962-38973 (12), stream at Le 

 Ninh, Quang Binh Prov., Vietnam, 17°14.12'N 

 106°37.51'E, 8 Apr 2000, J. Freyhof & F Herder; ZFMK 

 35462-36465 (4), stream at Le Ninh, Quang Binh Prov., 

 Vietnam, 17°13.16'N 106°39.08'E, 8 Apr 2000, J. Frey- 

 hof & F. Herder. 



Remarks: Winstanley & Clements (2008) were not able 

 to follow unambiguously the diagnostic colour characters 

 used by Freyhof & Herder (2002) for discriminating M. 

 erythropterus from M. spechti, and suggested that M. ery- 

 thropterus constitutes a junior synonym ofM. spechti. As 

 their analyses did however (i) not incorporate material 

 from the terra typica (streams Quyet Tien and Le Ninh, 

 Quang Binh Province, Vietnam), (ii) were based on only 

 limited numbers of obviously mostly immature specimens 

 (N totaI = 22 ex., without reference to maturity or sex; 

 SL max = 42.3 mm SL; four samling locations incorporat- 

 ed: AK 27598, 6 ex.; 1 3.7-38.0 mm SL; AK 27599, 3 ex., 

 19.1-26.0 mm SL; AK 27600, 3 ex., 26.0-34.1 mm SL; 

 AK 27601, 10 ex., 26.5-^12.3 mm SL; compare the sizes 

 of specimens examined by Freyhof & Herder (2002) as 

 listed above), and (iii) covered only locations from a part 

 of the species distribution (only locations in the Quang Tri 

 Province in Vietnam), we argue that this conclusion is pre- 

 mature. Macropodus erythropterus is, like many other 

 teleost fish species, distinguished from other species of its 

 genus mainly by colour characters present in adult spec- 

 imens. As long as these are not incorporated properly in 

 critical analyses, we consider the database for judging 

 about species status insufficient. Hence, we suggest that 

 M. erythropterus should be treated as valid species. 



Parosphromenus nagyi Schaller, 1985 

 Aquar Terrar Z 38: 302 



Lectotype: ZFMK 14186, Asian highway route 18, 16 km 

 south of Kuantan, Malaysia, 22 Feb 1985, D. Schaller. 



Paralectotypes: ZFMK 14187-14191 (5), same data as lec- 

 totype. 



©ZFMK 



