55 



APPENDIX. 



Page 15. 



Garryace^;. 



Fadgenia laurifolia has the ovary and ovules of a Cornus supposing the dissepiment in the latter to be removed ; — in all the flowers examined the dissepiment if present would pass between 

 the ovules, as the stigmas are always opposite them ; — the thick, twisted raphe and funiculus, like that of C. sanguined, may be seen turning up in opposite directions in the two ovules, and 

 the canal in the style in most instances opens over the outside of each ovule. The difference in the wood, more perhaps than any other character, requires this family to be placed in the Apetalous 

 Division, because in the absence of the dissepiment they may be well compared with Vahlia among Saxifragacece. 



Page 35. 



Orobanchace^e. 



The two carpels in Lathrcea squamaria are anterior and posterior, which is clearly proved by the placentae being right and left of the axis, and the stigmas anterior and posterior in the expanded 

 flower, and also in the unopened flower when so young that scarcely any style is formed ; — in the expanded flower the anterior lobe is larger as in some genera of Acanthaeece. Each placenta 

 however separates partially into two as it ascends, so far at least as to become naked and less prominent in the middle. As Orobanche is so near Lathrcea, differing more in the stigmas being 

 right and left than in any other character, it follows that the placentation in that genus is not dorsal because it is the same as that of Lathrcea, the separation in each placenta being rather wider 

 but yet not extending half-way from the margin towards the mid-rib of the carpel. In all other instances of dorsal placentation I have examined there is no placenta formed, the ovules arising 

 from the substance of the carpel, but in Orobanche as in Lathrcea two very thick placentae are present, but they divide as they ascend into four, the halves thus separated however closely 

 approximating again at the apex of the ovary. This being the case, the apparent dorsal placentation of Orobanchacece may be entirely owing to a displacement of the placentae as in 

 Mesembryanthemum, which is the more probable as it occurs only in a part of the genera. The stigmas being right and left in Orobanche is accounted for by the early condition of the flower in 

 O. major and elatior, and especially the latter, which might be described as having the stigmas anterior and posterior as in Lathrcea, but by the time the flower has expanded two lateral lobes have 

 grown and obliterated this appearance, but the tranverse fissure remains in the same direction in both genera. In Acanthus spinosa the two carpels are anterior and posterior, but the stigmas 

 right and left, which would perhaps be sufficient to account for this deviation in Orobanche. 



Page 37* 

 AcANTHACEiE. 



The radicle in Thunbergiece is described as inferior, but this seems impossible as the foramen in Thunbergia has distinctly the appearance of being superior, and the inferior portion of the 

 secundine is broadly oval or almost flat ; and Prof. Agardh, who I have always found correct as to the position of the foramen as far as regards its being superior or inferior, states that it is superior 

 (Theor. Syst. Plant., p. 189), which is also seen in his figure of T. alata, PI. XVII., fig. 2. The probability appears to be that the ovule curves more or less as it advances in growth, so that 

 the radicle is directed to the hilum, and is therefore horizontal as in Spielmannia. v. Table III. Spielmannia has also been described as having an inferior radicle although the seed is pendulous, 

 being attached close to the apex of the cell, but the radicle is directed away from the hilum laterally, the embryo lying horizontally across the seed. 



