NUPHAR INTERMEDIA LEDEB., A PRESUMED RELICT HYBRID, IN 



BRITAIN 



By YoLANDE Heslop-Harrison ✓ 



University College, London 



Introduction 



Nuphar intermedia was originally described by Ledebour (1832) from plants obtained 

 from Heiligenzee and Zee von Camby in North Germany. As was implied by the name 

 given it, the species was intended to embrace plants which lay between the generally 

 accepted variation ranges of N. lutea (L.) Sm. and N. pumila (Timm) DC. Subsequently, 

 Caspary (1870) and other students of the Nymphaeaceae pointed out the similarity between 

 wild plants which had been referred to Ledebour' s N. intermedia and artificial hybrids 



Fig. 1. Scottish and northern Enghsh ranges of N. mtcnncdia and it^ presumed parents N. pwnila and 

 N. lutca. N. intermedia occurs at open circles, N. pwnila at black circles and N. lutca in shaded vicc-counties. 



7 



