450 Miss Tozer and Prof. Sherrington, Receptors and [May 18, 



Morris with attached than with detached leaves of H. annuus cannot be 

 entirely attributed to less widely opened stomata ; but that part of the 

 products of photosynthesis must have been translocated. 



Broocks' experiments with the sugar-beet have proved that in that plant 

 translocation does proceed during the day. On two continuously sunny days 

 he found that leaves increased rapidly in dry weight till noon, and then, 

 suddenly ceasing to assimilate, lost weight continuously at a uniform rate 

 till about midnight. In this case also it seems more reasonable to suppose 

 that in the morning translocation was taking place, though masked by 

 assimilation, rather than that it only began when the leaves stopped 

 assimilating. 



I hope myself to obtain more conchtsive evidence as to whether assimila- 

 tion and translocation proceed concurrently. Now that a satisfactory 

 modification of the dry weight method has been elaborated it is to be expected 

 that this and other similar problems will be successfully attacked. 



Receptors mid Afferents of the Third, Fourth, and Sixth Cranial 



Nerves. 



By Frances M. Tozer, B.Sc, and C. S. Sherrington, D.Sc. 

 (Received May 18,— Read June 6, 1910.) 



The view generally accepted regarding the functions of the third, fourth, 

 and sixth cranial nerve pairs is that they are purely motor. Certain experi- 

 ments and observations made by one of us* and published some years ago 

 tlirew doubt, however, on this belief. 



It was then shown that severance of the third or fourth or sixth nerves at 

 origin from the brain produces degeneration of practically all the nerve- 

 fibres of the respective muscles innervated by those nerves and of the 

 receptive endorgans with which those muscles are, as was shown.f plentifully 

 supplied. 



Further, on the ])liysiological side it was found (1) tliat intracranial 

 severance of both nn. trigemini in the monkey resulted in no obvious 

 impairment or ataxy of eyeball movements, and (2) that severance of both 



* ' Physiol Soc. Proc.,' June, 1894 ; ' Journ. of Physiol.,' vol. 17, p. 20 ; ' Roy. Soc. Proc.,' 

 Feb., 1897, vol. 61, p. 247 ; ibid., Sept., 1898, vol. 64, p. 120. 

 + Sherrington, 'Roy. Soc. Proc.,' Veh., 1897. 



