TOMATO DISEASE IN GUERNSEY. 
28 
was the case with the two previous stages. Fig. 3, a, shows 
the formation of chlamydospores on the hyphre ; b, a mature 
free chlamydospore ; c, the same germinating, and producing 
the first or Diplocladium form of fruit. The resting-spores, as 
before stated, are present in the soil, and attack the roots of 
young Tomatos ; hence the importance of removing all diseased 
plants before the resting-spores are formed, as these pass into 
the soil when the Tomato roots, where they are principally 
formed, decay. It is not sufficient to simply pull up the diseased 
plants, but all the rootlets should be removed, as these are the 
parts first attacked by the fungus, and consequently contain the 
most matured hyphre, and, being surrounded by soil, the usual 
conidial stages are not produced; consequently the resting-spores 
are developed in greater quantities than in the above-ground 
parts of the plant. 
It is somewhat remarkable, and contrary to the general rule, 
that the conidia of the rapidly maturing summer phases of the 
fungus — Diplocladium and Fusarium — whose function is to effect 
the spread of the fungus, should be unable to attack the living 
tissues of the Tomato, but such is the case ; the conidia of both 
forms germinate only as saprophytes, growing readily on rich 
humus, manure, &c., the conidia of the Diplocladium on germi- 
nation producing the Fusarium, the hypha) finally forming 
resting-spores in the soil, the Fusarium conidia giving origin to 
its like and resting-spores onthehyphae. Thus both stages, after 
extending their area during the summer months, agree in pro- 
ducing resting-spores in the soil, which renew the cycle of 
development by germinating and attacking young Tomatos during 
the spring following their formation. 
Fusarium Lycopersici was first described by Saccardo in 1882, 
from Italian specimens, growing on decaying Tomatos, but was 
not at that time connected with a disease. 
Summary. 
From what has been said, it will have been gathered that the 
germinating resting- spore is the only condition of the fungus 
capable of attacking the Tomato ; hence the characteristics of 
the disease — drooping of the leaves in succession from the base 
of the plant upwards, and the discoloration of the wood in the 
root — should be thoroughly grasped, and plants promptly removed 
