276 



MONOGRAPH OF THE LABOULBEXIACE M. 



This curious species occurs rather rarely, more often on the upper than on the inferior sur- 

 face of the abdomen of the two hosts mentioned, the receptacle lying close against the surface, 

 while the perithecia project outward, often backward. Specimens on S. longiusculus are 

 somewhat smaller and much paler than those on the larger host, the basal cell of the receptacle 

 being more or less translucent. The form is not easily overlooked when mature, occurring usu- 

 ally in groups of half-a-dozen or more. The hosts may be found in dry hay or other rubbish in 

 cultivated land. 



ENARTHROMYCES nov. gen. Plates III-IV. 



Receptacle consisting of a simple series of superposed cells, the distal ones bearing sterile 

 appendages, those below giving rise to antheridia or perithecia or remaining sterile. Anthe- 

 ridia compound, terminally pointed and perforate, the six antheridial cells converging upward 

 to a general cavity, into which they empty through narrow necks. Trichogyne simple, two- 

 celled. Perithecia stalked, one or more formed by direct budding from the cells of the 

 receptacle. 



In the form and structure of its antheridium, and to some extent of its perithecium, this re- 

 markable genus recalls Peyritschiella, to which it is undoubtedly more closely related than to 

 any other. The series of specimens which constitute the types of the single species contain no 

 very young individuals ; yet it is apparent that, after the axis is formed and most of the sterile 

 appendages have been produced, an antheridium arises, first originating as a lateral bud, in 

 such a position that the first trichogyne when mature lies beside its apex. The antheridium is 

 the largest thus far observed, of the compound type, and its general structure may be made out 

 with no great difficulty. The antheridial cells are irregularly flask-shaped, and occupy the 

 somewhat inflated basal portion of the organ, the distal part being more or less conical, and 

 apparently originating from a single cell, the remains of a septum being usually visible (Plate 

 III, fig. 19) just above the openings of the canals through which the contents of the antheridial 

 cells are discharged. The cavity which it contains is of considerable size, and is often filled 

 with free antherozoids. The latter are larger than other known forms of endogenous origin, and 

 seem to be surrounded by a thin wall at the time of their final discharge. The trichogyne is 

 quite peculiar, and invariable in form, being rather short and inflated, septate near the middle, 

 and prolonged into a short terminal projection (Plate III, fig. 17, tr), which seems to be the 

 only portion that is receptive, the remainder being distinctly thick walled. Indeed, it is only 

 the terminal part of this projection that is very thin walled ; so that, after fertilization, when this 

 receptive portion has withered, the trichogyne appears to end in a short, cylindrical, and distally 

 truncate and perforate outgrowth (fig. 18, t r). The young perithecia arise as lateral buds from 

 the cells of the axis, becoming cut off from it in the form of free clavate cells (fig. 13), which 

 become once septate (fig. 14) and follow the course of development described in Part I (p. 228). 



Enarthromyces indicus nov. sp. Plate III, figs. 13-19 ; Plate IV, figs. 8-11. 



Pale amber or straw colored. Perithecia large, sub-cylindrical, terminally rather abruptly 

 and symmetrically rounded below the short, clearly distinguished, broadly truncate apex, the lip- 



