296 



MONOGRAPH OF THE LABOULBENIACEiE. 



that becomes separated from the end of the trichophoric cell, and corresponds to the indurated 

 portion of the trichogyne described in connection with Camptomyces. From this cell the recep- 

 tive portion of the trichogyne first makes its appearance as a minute papilla, which enlarges 

 and grows out to form the characteristic branches just described. 



The development of the perithecium proper has not been followed out in all its details, but 

 seems to correspond closely to that described as typical in the first part of this Memoir. 

 Whether there is a stalk-cell and a secondary stalk-cell has not been determined, but it is 

 probable that one of the two may be absent, since there seem to be but four cells about the base 

 of the perithecium. 



I have also been unable to observe the first divisions which take place in the carpogenic cell 

 after fertilization, and it is only after the asci have begun to bud from the ascogenic cell that 

 the interior of the perithecium can be seen with sufficient clearness to determine the nature of 

 the changes which are taking place. At this period, however, the ascogenic cell becomes very 

 distinct, and from its large size is a very good subject for observation. Although it is very long 

 and there is sometimes an indication of a septum across it, its development appears to be iden- 

 tical with that of other cases already described. Its period of activity does not seem, despite 

 its unusual size, to be as great as that of most ascogenic cells ; and it is in this genus only that 

 one is very apt to find individuals which have apparently died after completing their period of 

 natural activity. 



As in Dimorphomyces, instances sometimes occur in which the spores, not having been dis- 

 charged for some reason, begin to germinate within the perithecium ; and specimens like that 

 represented in fig. 19 are not of very rare occurrence, the blackened foot in such instances being 

 conspicuously developed, as well as faint indications of the three primary septa; but no case of 

 almost complete development like that mentioned in Dimorphomyces (fig. 1) has been observed 

 in the present instance. 



The affinities of this very important genus are very obscure. From the character of its 

 antheridium it would be placed in a different group from Dimorphomyces, which occurs on the 

 same host and possesses an antheridium of the compound type very highly developed ; yet in 

 some respects it seems to approach this genus as closely as any other. The two genera are 

 often found together on the same host, which is a very small beetle belonging to the 

 Staphylinidae. 



Amorphomyces Falagri^e Thaxter. Plate V, figs. 17-29. 



Proc. Am. Acad. Arts and Sci. Vol. XXVIII, p. 158. 



Male individual consisting of three superposed cells, the basal and sub-basal about equal, the 

 latter suffused with dark brown, the suffusion often extending to the adjacent cells ; the distal 

 cell an antheridium, basally slightly inflated, distally prolonged into a cylindrical neck bent to 

 one side. Total length, 48 /m by 10 n broad. 



Female individual. Receptacle hyaline, consisting of a small basal cell, sometimes partly 

 divided, surmounted by usually three small cells more or less irregular in shape and position, 

 which form the base of the large pale brownish yellow perithecium, which is curved strongly on 

 the side opposite the ascogenic cell, and tapers to a blunt almost truncate or abruptly lobed tip. 



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