250 JOURNAL OF THE ROYAL HORTICULTURAL SOCIETY. 
work with respect to false crossing, a series of new non- separating crosses.* 
These he obtains by union of Fragaria, Buhus, &c., and found them to be 
uniform in the first generation ; they resembled either the father or the 
mother. He proposed for the non- separating Hybrids the name of False 
Crosses ; consequently those cases in which Mendel's law of separation is 
followed may be termed True Crosses. 
With these two cases before us there exists obviously a third possi- 
bility, viz., that crosses may separate according to other rules. Such 
cases, indeed, I have found to exist ; they showed themselves to be far 
more nearly related to false crosses than to true ones, and are therefore 
ranked with the former. 
I will therefore henceforth name those crossings whose produce, when 
the sexual organs are formed, separates equally, and which therefore 
follow Mendel's law, Equal Heritors (erbgleich) or Isogons, and their 
progeny — i.e. the crosses themselves — True Crosses. 
On the other hand I will call those crossings whose produce at such 
stage either does not separate, or separates according to other rules, Unequal 
Heritors {erhtngleich) or Anisogons, and their progeny, as Millardet does. 
False Crosses. 
There appears at present no reason to assume that the crossings of 
unequal heritage in the plant world are rarer than those of equal 
heritage,t and the very general occurrence of the latter naturally does not 
stand in the way of this. So far as my experience has hitherto gone, the 
former are certainly no less numerous than the latter. In addition to 
the genus CEnothcra, of which I intend here to treat, I found several 
characters of unequal heritage which are widely spread in the plant 
world, as, for instance, Polycephaly, Tricotyly, Syncotyly, &c. On the 
whole 1 think that I have made far more crossings of unequal heritage 
than of equal, but this can hardly be numerically defined. 
In my previous paper on Separation of Characters I expressly con- 
fined myself to the true crosses, and for the time being wholly disregarded 
the false hybrids of Millardet (see note, p. 244). I have there deferred the 
consideration of these for another paper, and purpose now to give a pre- 
liminary report upon them. I will, however, only refer to my crossings 
in the genus CEnotliera.X 
(Enothera muricata $ x biennis ^ § is a typical false cross, which 
agrees exactly with the examples given by Millardet. It displays the 
paternal character with the exception of fertility, which is much reduced, 
while, in accordance therewith, the vegetative parts are more luxuriantly 
developed. The stalks, and especially the flower stems, are far more 
richly furnished with leaves. This cross I effected in 1895. In the first 
generation I had fifty crossed progeny, all like each other. I collected 
the seed from two-year-old examples (1897) and have since had three 
other one-year-old generations of about same extent. Separation did not 
* A. Millardet, -'Note sur I'Hybridation sans Croisement, ou Fausse Hybridation," 
M6m. Soc. Sc. Pliys. et Nat. Bordeaux, vol. iv. (4 Series, 1894, pp. 1-28). 
f Correns is certainly of a different opinion, and regards the cited Hieraciuvi 
crosses as exceptions to Mendel's law. 
J For the description of the new species of this genus named hereafter, I refer to 
the first volume of my Miitationstheorie (Leipzig : Veit & Co.) ; and for detailed 
description of my crossing experiments, to the second volume of the same work. 
§ Compare Focke, Die Pflanzenmischlinge, p. 163. 
