Xlii PROCEEDINGS OF THE ROYAL HORTICULTURAL SOCIETY. 
of an orange colour, but as usual in staminate inflorescences of this plant, 
containing no seeds. 
Flowering of Lilium philippinense.' — Mrs. Brown also referred to the rapidity 
with which seedlings of L. philippinense reach flowering size. Plants raised 
from seed flowered within fifteen months of the germination of the seeds. 
Various Plants. — Mr. H.J. Elwes showed a number of plants from his garden, 
including Campanula Vidalii, a rather tender species with a stem somewhat 
woody at the base, an unnamed species of Allium collected by Captain Bailey 
in Tibet, which spreads by means of rhizomes, and is related to A . W allichianum ; 
Urceocharis Clibranii which is said to be of hybrid origin, but which bears a 
somewhat close resemblance to a plant sent home from South America by 
Forget ; Roscoea Humeana, a Chinese species which Mr. Elwes regards as a 
finer plant than R. purpurea ; and Phlomis spectabilis, an uncommon species. 
Scientific Committee, September 7, 1920. 
Mr. E. A. Bowles, M.A., in the Chair, two members and Dr. R. Gates present. 
Starch Roots in Oxalis. — Sir Archibald Hepburn sent a tuber of Oxalis adeno- 
phylla with a starch root such as is formed in many species of Oxalis. 
Hybrid Oenotheras . — Dr. R. R. Gates showed a series of hybrid Oenotheras 
and commented upon them as follows : This is a series of hybrids between 
Oenothera rubricalyx and certain wild species. 0. rubricalyx is distinguished 
by having deep red instead of green buds. A single plant of this type appeared 
in my experimental cultures as a mutation in 1907, and all red-budded Oenotheras 
in cultivation are, without exception, descended from this plant and various 
crosses subsequently made with its descendants. O. rubricalyx was a mutation 
from O. rubrinervis, which in turn was derived in the cultures of de Vries from 
O. Lamar ckiana. In its offspring this rubricalyx plant behaved at first as a 
simple Mendelian dominant, giving 3 red-budded to i green-budded offspring. 
In later generations a complication was introduced in the appearance of 15:1 
ratios in addition to 3 : i, indicating the appearance of a second factor for red. 
A pure race was also obtained by inbreeding, and in 191 2 Messrs. Sutton acquired 
seeds of it (partly crossed with O. grandiflora) and introduced it into horticulture 
under the name ' Afterglow.' 
Although evening Primroses have been in cultivation since 161 4, this orna- 
mental red variety has appeared but once, showing the great rarity of this 
mutation. In crosses this character of red buds always behaves as a dominant 
though differing somewhat in the details of its behaviour in species crosses. 
Among the numerous hybrids with other species and varieties which have been 
made in my experiments, the following were shown : — 
(1) 0. Hewettii x O. rubricalyx, Fg. 
O. Hewettii from Colorado is an exceptionally tall species with large flowers. 
This hybrid has very good horticultural characters, and an Fg generation will 
be grown for further study and to select a race which is pure for dark-red buds. 
(2) 0. rubricalyx x O. Hewettii, Fj. 
The buds are a clear red, and the plants have a fine pyramidal habit of 
branching. 
(3) O. Novae Scotiae x O. rubricalyx, Fj. 
O. Novae Scotiae (Gates) was recently described from Nova Scotia. It has 
very small flowers and in the F^ hybrids the size of petal is intermediate.- The 
red buds, however, give a very ornamental effect, and the Fg generation will 
no doubt yield plants with larger flowers. 
(4) O. rubricalyx x O. biennis, F4. 
A number of families from this cross, inbred for three generations, 
yield uniform offspring with dark red buds, differing somewhat in size of flower. 
(5) Various hybrids of rubricalyx with green-budded forms give races with 
uniformly pale-red buds. 
Scientific Committee, September 21, 1920. 
. Mr. E. A. Bowles, M.A., in the Chair, with four members present. 
Corm 0} Cyclamen. — Mr. H.J. Chapman sent a corm of Cyclamen neapolitanum 
to illustrate the large size these corms attain. It was planted forty years ago, 
and had grown and flowered well ever since, and now measured 8 inches in 
diameter. 
