230 



Mycologia 



Illustrations : Giesenhagen, Ber. Deut. Bot. Gesell. 22 : pi. 

 13. 1904; Stevens, 111. Acad. Sci. Trans. 10; fig. 6. 19 17. 



Type : in the herbarium of Berkeley, at the Royal Botanic 

 Gardens, Kew, England ; co-type material in the herbarium of 

 Curtis at Harvard University. Both of these have been ex- 

 amined by the writer, and a portion of the type is deposited in 

 his herbarium. 



(Figures 30-32, 49) y 

 Mycelium parasitic in the leaves of the host, localized, confined 

 to a small area surrounding the point of infection, forming a 

 definite stroma within the host tissue ; stromata in some collec- 

 tions of material scattered irregularly over the leaf, in others 

 confined to the sori, the general appearance of the fungus on the 

 host in the two cases consequently very different ; the stroma 

 when formed in the sorus not well-developed, buried from sight 

 below the mass of host sporangia, not spreading to the other 

 host tissues, usually not forming a spot in the leaf, and never 

 observed to result in its perforation ; perithecia arising as black 

 bristles among the host sporangia and radiating in every direc- 

 tion ; stromata when formed in other portions of the leaf erum- 

 pent, usually hypophyllous, at first minute, gradually increasing 

 in area, the central portion soon falling out, leaving only the 

 opposite epidermis or more often a complete perforation ; the 

 hole thus formed in the leaf reaching 3-4 mm. in diameter, sur- 

 rounded by an annular or ring-shaped stroma, less than 1 mm. in 

 width, and of the thickness of the leaf or slightly thicker ; 

 stromata from the first bearing numerous pycnidia and peri- 

 thecia ; later when the hole appears in the 'leaf the perithecia 

 forming a ring of bristles around the perforation and arising 

 from the annular stroma on both the upper and lower surfaces ; 

 perithecium shiny to dull, 1— 1 .5 mm. long, stalk 35-50//, thick, the 

 ascigerous portion glabrous, 125-150^ in diam. X 250^ in length, 

 tapering into a glabrous beak-like neck, 200-350 fx long; stalk 

 frequently hairy with brown hyphae ; asci 15-17 X 10 jjl (p. sp.), 

 ascospores 5-6 X 4-5 ^ ; pycnidia covered with brown hairs like 

 those on the stalk of the perithecium ; pycnosoores 11-24X4^. 



Parasitic on Poly podium (Campyloneurum<) phyllitidis, P. 

 punctatum, P. crassifolium, P '. Schomburghianiim' and probably 

 other species in Cuba, Porto Rico, San Domingo, Brazil, Ecua- 

 dor, Venezuela, and probably other neighboring countries. 



Giesenhagen (15) states that the brown hairs which clothe the 

 pycnidium and the stalk of the perithecium are conidial bearing 



