Shear and Dodge : Patellina, Leptothyrium, Peziza137 



on blackberries in New Jersey. A few sporodochia of the same 

 fungus were found in April, 1920, on old dewberry vines from 

 Cameron, North Carolina, and cultures were readily obtained 

 from this material, showing that the fungus had survived the 

 winter on the old vines. 



The material from North Carolina was also found to bear con- 

 spicuous brown or black sclerotium-like pycnidia which proved 

 to be Leptothyrium macro the cium Fckl. This was reported by 

 Fuckel to occur on a variety of hosts, one of which was Rubus. 

 On account of the close association of the Hainesia sporodochia 

 and the Leptothyrium pycnidia and the great similarity of the 

 spores of the two forms a possible genetic relation was suspected. 

 If this was true the Leptothyrium stage, being evidently of a 

 more persistent and resistant character than the other, might be 

 the means of carrying the fungus over the winter. It may be noted 

 here that cultures from these pyenospores produced typical coni- 

 dial sporodochia. The pycnidial stage was also produced at will 

 on leaves and stems of Rubus and other plants by spraying with a 

 suspension of conidia. 



xA.ssuming that this fungus had but two lower spore forms it 

 now remained to find the ascogenous stage. If any similarity in 

 form was to be expected between the pyenidium and the ascocarp, 

 Hypoderma might perhaps be suggested on account of its slight 

 superficial resemblance to the pycnidia and its occurrence on some 

 of the dewberry canes. On the other hand the sporodochia sug- 

 gest in form a possible small discomycete of similar appearance. 

 The discovery a little later of a small amber-colored discomycete 

 on old leaves of raspberry at Arlington Farm, Virginia, July 24, 

 closely associated with both sporodochia and pycnidia was imme- 

 diately followed by pure cultures from ascospores which proved 

 the genetic relation of the three forms. A search of literature 

 and herbaria showed that a discomycete apparently agreeing in 

 all respects with the one found on raspberry leaves, had been 

 described as Peziza (Mollisia) oenotherae C. & E. (1878) and 

 distributed as No. 846, Ellis and Everhart, N. A. F. and 244 Fun. 

 Col. All three forms were found occurring together on stems of 

 Oenothera biennis on the same herbarium specimen of No. 244 



