118 



Mycologia 



threads were clearly seen, and their exact position in regard to 

 the asci was shown. The presence of the threads between the 

 asci was demonstrated with a certainty and can be clearly seen 

 in the accompanying photomicrographs. While I have made 

 scores of slides from other host plants, I was never before able 

 to demonstrate their presence between the asci, and in a pre- 

 vious paper I expressed a doubt as to their occurrence in that 

 position. The reason evidently lay in the better development 

 of the threads in the perithecia from the cotton. These threads 

 are much longer than the asci, extending to the ostiole of the 

 perithecium, and entirely filling the cavity above the asci. If 

 these sterile threads are to be called paraphyses, and there is 

 little reason why they should not be called so, notwithstanding 

 their length and irregularity, then the genus Glomerella must in 

 the future be considered as paraphysate. 



What effect this may have on retaining or discarding the gen- 

 eric name, Glomerella, cannot be told until some one has made a 

 careful study of the various species of the genus Physalospora. 

 Some of the perithecia of the cotton anthracnose fit perfectly the 

 characters of the genus Physalospora, as shown in fig. 3, of the 

 accompanying plate ; however, others, as in fig. 2, do not. Glom- 

 erella perithecia, on artificial media or in a moist chamber, gen- 

 erally develop on the surface of the substratum and are not 

 embedded in it, while Physalospora perithecia are not supposed 

 to develop in this manner. But if it is found that members of 

 this latter genus will develop on the surface of the substratum 

 if the moisture conditions are suitable, then there are no generic 

 characters separating the forms now resting in- Glomerella and 

 Physalospora. Several mycologists, such as Maublanc and Las- 

 nier in France, and Sheldon in this country, are now calling the 

 anthracnose forms Physalospora; but, until we know more of 

 the development of these different forms, it seems best to keep 

 them separate. • 



Although many of the anthracnoses found on different hosts 

 are members of a single species, as has been shown by various 

 investigators in the last few years, the evidence seems to show that 

 some forms are distinct enough to represent different species. 

 While morphologically the different forms are in many ways 



