236 



Mycologia 



mens studied, and the similar effect produced on the algal host 

 by the two lichens, together with their similarity with respect to 

 apothecia and spores, led to the suspicion that the plants might 

 after all belong to the same species, the internal spermatia being 

 conidia, with which the trichogynes fused in some instances in- 

 stead of with the spermatia produced in spermagonia. Though 

 this possibility was kept in mind in the examination of material, 

 in no instance were spermagonia and the free internal spermatia 

 found on the same plants, or in plants belonging to the same 

 collection. 



Before studying the Ohio material, it had been ascertained that 

 the material widely distributed by the writer from Fayette, Iowa, 

 in 1894 and following years, as Collema pulposum (Bernh.) Ach. 

 was not that lichen but the new and peculiar one studied first by 

 Miss Bachman and described in the present paper. After work- 

 ing on the Ohio material, the specimens in the writer's herbarium 

 were gone over with the result that the new plant was found 

 from widely separate stations, extending from the Atlantic coast 

 to areas west of the Mississippi River. 



On account of the similar modification of the algal-host colony 

 and the likeness with respect to apothecia and spores between the 

 plant described below and certain species of Collema, especially 

 Collema pulposum (Bernh.) Ach., one can never hope to make 

 sure whether he has our plant or a Collema, without ascertaining 

 the morphology of the male reproductive tract, and the behavior 

 of both male and female reproductive tracts. In these respects 

 it must be remembered that our plant has internal, much larger 

 spermatia, not contained in spermagonia and that the trichogynes 

 grow to the spermatia within the thallus ; while in the Collemae 

 the trichogyne extends above the surface of the thallus, and the 

 much smaller spermatia escape from the superficial spermagonia 

 and are carried to this exposed portion of the female reproduc- 

 tive tract. 



Collemodes gen. nov. 



Transforming the algal-host colonies into f oliose bodies ; thallus 

 wholly mycelial, imbedded in the host colonies, and attached to 

 the substratum by rhizoids ; male reproductive organs not in 



