246 
PHYSIOLOGY: ATKINSON AND LUSK 
times as much to the weight of k as do those in zone II. The mean result 
for k should therefore be sensibly the same as the values for zones I and III 
when treated separately, as proves to be the case (k = 0.99, 96, and 96, re- 
spectively). The weight of the inclination, on the other hand, is derived 
largely from the observations of the equatorial zone. Moreover, i is calcu- 
lated after k has been found, so that the conditions in the uniform-field solu- 
tion are analogous to those of the first approximation of zone II. In this 
approximation the equatorial observations combined with x'= 1.00 — the equiv- 
alent of ^ = 0.99 — gave i = 6?7, which is nearly the same as the value found 
from all the data. The measures of zones I and III combined with k = 0.99, 
as we know from the zonal analysis, must lead to relatively small values for 
i, amounting to about 4°. The mean therefore lies between these two ex- 
tremes, with a preponderance of weight in zone II. This accounts for the 
value of 6?0 originally found — a mean result which is in excess of all the in- 
clinations found by treating the zones separately. It is only when the equa- 
torial observations are discussed by themselves that the true value of k for this 
region reveals itself or has any appreciable influence upon the solution; but 
when once found the inclination is necessarily decreased. 
We are under great obligation to Miss Wolfe of the Computing Division 
who has rendered much assistance with the extensive numerical calculations 
required for the discussion of the data. 
1 Scares, F. H., van Maanen, A., and Ellerman, F., these Proceedings 4, 1918, (A-^)). 
ON THE PROBLEM OF THE PRODUCTION OF FAT FROM PROTEIN 
IN THE DOG 
By H. V. Atkinson and Graham Lusk 
Cornell University Medical College, New York City 
Read before the Academy, April 28, 1919 
In eight experiments in a series of thirteen, after giving meat in large quan- 
tities (700 to 1300 grams) to a dog weighing 11 kgm., the respiratory quotients 
during the height of protein metabolism were between 0.793 and 0.800. A 
calculation showed that under these conditions the retained carbon residue 
of the protein metabolized was held back in such a form that, had it been 
oxidized, the respiratory quotient of this retained pabulum would have been 
0.85. This would represent the oxidation of material half of whose calories 
were composed of fat and half of carbohydrate. The dog showed quotients 
of 0.82 and above only after the larger quantities of meat were given (1000 
grams or more). It was extremely difficult to induce the dog to take meat in 
sufficient quantity to indicate a considerable production of fat from protein. 
Incidentally it was observed that the basal metabolism of a dog fed with meat 
in large quantity for a time and then caused to revert to a 'standard diet' 
