ZOOLOGY: KOFOID AND SWEZY 
319 
mose later fades out after the karyosomes and chromatin network re- 
appear within the daughter nuclei (fig. 7). 
At the close of the telophase (fig. 7) division of the axostyle proceeds 
by longitudinal spHtting from the anterior end posteriorly. There is a 
suggestion that axostylar chromidia are also divided, at least in the dis- 
tal region (fig. 7). Actual division of individual chromidia has not 
been seen though in some dividing axostyles (fig. 7) their relative dimen- 
sions in parent and daughter axostyles strongly suggest their division. 
The numbers of chromidia vary so that they do not afford critical evi- 
dence for use on this point. 
The proofs of division of the axostyle in our material are ample though 
their detection requires well-prepared slides and persistent search. 
Dobell^ and Prowazek^ describe the origin of these axostyles from the 
*'centrodesmose" and the former concludes that it is therefore the homo- 
logue of the central spindle, and Doflein in his recent Lehrbucher der Pro- 
tozoenkunde accepts this origin. This strand throughout all our material 
is everywhere extra-nuclear and we therefore propose for it the name 
paradesmose since it cannot occupy the position of the central spindle 
or "centrodesmose.'^ Kuczynski^ has shown that DobelFs and Pro- 
wazek's conclusions are invaHd since the paradesmose persists till after 
the new axostyles appear. This author, however, falls into the error of 
concluding that the daughter axostyles arise as new outgrowths from 
certain end granules of the old axostyle originally connected with the 
blepharoplast. He finds that the old axostyle fades out and the daughter 
axostyles arise as new outgrowths. In our material it has always been 
possible to detect in all individuals which have passed the anaphase of 
mitosis one of three conditions, either (1) the single undivided axostyle 
or (2) this axostyle in process of splitting longitudinally, or (3) two 
complete daughter axostyles. The presence of the axostylar chromidia 
has been of great assistance in following the longitudinal splitting of 
this organ in Trichomonas augusta. A similar splitting of the axostyle 
occurs in Lamhlia muris and in two or more species of Hexamitus. 
Wenyon^ states that the axostyle of Trichomonas intestinalis of the 
mouse divides longitudinally after the division of the nucleus and 
blepharoplast. His figure, however, does not support his statement. 
It shows two axostyles rather than a dividing one. 
Shortly after the axostyle splits the two nuclei and axostyles move 
farther apart, the body elongates, the anterior ends of the two daughters 
come to lie at opposite poles (fig. 7) and the cytoplasmic central bridge 
progressively narrows down until the two daughter organisms separate. 
The final plane of constriction is transverse to the main axis of each cell 
and of the common elongating mass. 
