102 THOMPSON YATES AND JOHNSTON LABORATORIES REPORT 
therefore amounts to eight. In the paper already referred to Dr. Brumpt considers 
that Trpanosoma brucei, the parasite of nagana or Tsetse-fly disease in domestic animals, 
is carried by at least five of these species/ and he further states that the investigations 
that he is making upon sleeping sleepness lead him to suppose that this malady may 
also be transmitted by several species of Tsetse-flies. The mere possibility that this 
supposition may ultimately prove to be true is perhaps sufficient warrant for thinking 
that no detail concerning the morphological characters, distribution, or bionomics of 
any of the species of Glossina is without importance for those interested in the sanita- 
tion of tropical Africa. In the present paper, therefore, the eight species will be 
considered in order, and any new facts regarded as worthy of notice recorded under 
each, the arrangement adopted in the Monograph being adhered to, and Glossina 
tacbinoldes inserted in its proper place. At the end of the paper a revised ' Synopsis 
of species ' will be given, which it is hoped may prove useful for the determination of 
specimens. 
Glossina palpalis (Rob.-Desv.) 
The form of this species designated in the Monograph as ' Var. tachinoides 
(Westw.) ' must now be regarded as a variety which for the present may remain un- 
named. It is characterized by the possession of pale femora, buff-yellow median stripe 
on the abdomen, and narrow pale hind margins to the abdominal segments. Specimens 
obtained by Drs. Todd and Dutton on the Gambia during the Gambia expedition of 
the Liverpool School of Tropical Medicine belong to this form, which may eventually 
have to be raised to specific rank. A second variety also has the femora paler than 
usual ; the palpi, except the tips, pale ; and the abdomen somewhat reddish-brown, 
with the pale area on the second segment oblong instead of triangular. Owing to the 
colour of the abdomen this form presents a certain resemblance to Glossina pallicera 
(Bigot). A specimen of variety No. 2 was obtained at Old Calabar on May 14, 1900, 
by Dr. Annett, of the Liverpool School of Tropical Medicine. 
As many of the members of this Section are doubtless aware, our knowledge of 
the life-history of Tsetse-flies was originally due to Colonel Bruce, who, during his 
investigations upon Tsetse-fly disease, or nagana, among domestic animals in Zululand 
in 1895, discovered that the species studied by him — either Glossina morsitans (Westw.) 
or Glossina pallidipes (Austen) — ' does not lay eggs as do the majority of Diptera, but 
extrudes a yellow-coloured larva nearly as large as the abdomen of the mother.' 3 Speci- 
mens of the pupa of this species, kindly supplied by Colonel Bruce, were described 
and figured on pp. 26-28 of the author's Monograph. Owing to the kindness of 
Colonel Bruce, Captain E. D. W. Greig, I. M.S., and Dr. Nabarro, all of the Sleep- 
ing Sickness Commission, who forwarded specimens from Entebbe, Uganda, it is now 
possible to describe the larva and pupa of Glossina palpalis. 
