FILICINEM. 417 
different ages {b, nb). The young leaves have a circinate vernation, and are com- 
pletely invested by the stipules until the time when the petiole begins to elongate 
and the lamina to unroll itself. Each pair of stipules belonging to a petiole forms, 
as is shown in Fig. 2gi, A and ^, an anterior and a posterior chamber, which are 
separated by a longitudinal wall (commissure). In the posterior chamber lies the 
rolled-up leaf, to which the stipules actually belong, the two posterior wings of the 
stipules extending round it. The chamber formed by the anterior wings of the 
stipules encloses the group of young leaves. This is the arrangement in Angiopteris, 
and it appears from herbarium specimens to obtain also in Dana;a, and from 
Fig. 291 A.- — Vertical section of the stem of a young A7igiopte7-is evecta : above are the youngest leaves (b) still completely 
surrounded by the stipules ; st petiole of an unfolded leaf with its stipule 71b; in every case the cicatrix on the basal 
portion of the petiole ff, from which the upper portions have separated ; c c the commissures of the stipules in vertical section ; 
iu TV roots (natural size). 
drawings in Maratiia. Harting's representation of the stipules is quite erroneous. 
These peculiar stipules remain fresh and succulent not only during the life of the 
leaves but also after they have fallen, and adventitious buds may originate from 
them. 
The roots arise, as is shown in Fig. 291, A, in the tissue of the stem 
immediately below the growing-point. One arises apparently at the base of each, 
young leaf. They grow obliquely through the succulent parenchyma of the stem 
and of the older basal portions of the leaves, and finally reach the surface at a 
E e 
