MATROCLINIC INHERITANCE 
ing to draw any conclusions from it, that seeds of mut. dehilis were 
much less viable than those of the crosses into which this form entered 
as the pistillate parent. 
The progeny obtained from the cross f. typica X mut. hilonga 
showed the influence of the pollen parent in the large number of yellow 
plants, the same, as far as superficial observation could indicate, as 
the yellow twin that appears in progenies from self -pollinated mut. 
hilonga. In view of the fact that the crosses typica X hilonga and 
dehilis X hilonga both resulted in twin types, it is interesting that the 
third cross, semialta X hilonga, gave only plants of the maternal type, 
aside from a few which appear to be satisfactorily accounted for as 
derived from mutated gametes. 
It will be observed from a scrutiny of Table II that in the foregoing 
discussion we have tacitly assumed that the sporadic occurrence of 
types in cultures where they would not necessarily be expected to 
occur was to be ascribed to mutation. We have made no special 
comment, for example, on the few individuals of mut. dehilis that turned 
up in the progeny of semialta X typica, for the reason that mut. 
semialta always seems to produce some mutated gametes that give 
rise on fertilization to mut. dehilis. Furthermore, there is every ground 
for the belief, on evidence furnished by the matroclinic progeny of the 
cross dehilis X typica, that the few dehilis-yieXdmg eggs of semialta 
would give rise to mut. dehilis quite regardless of the source of the 
male gametes, provided, of course, that the latter were not derived 
from m.ut. hilonga, the one form of the series that, seems to give rise 
to more than one type of sperms. 
To return to the case of the cross semialta X hilonga, we have seen 
that mut. semialta always produces some dehilis-yieXdmg eggs, and 
that the cross dehilis X hilonga yields a progeny containing both 
parental types. Consequently we should expect that in the cross 
semialta X hilonga some of the mutated eggs would give rise to mut. 
dehilis and some to mut. hilonga, whereas in a progeny resulting from 
self-pollination or from one of the other crosses the mutated eggs would 
be represented by mut. dehilis alone. The results of the crosses realize 
this expectation. Although mut. semialta, whether self-pollinated or 
crossed, has always given rise to mut. dehilis, it has never given mut. 
hilonga except in the case of the cross semialta X hilonga. 
The results of the whole series of cultures are intelligible on the 
supposition that Oe. Reynoldsii is one of the mutable species to which 
