402 
ORLAND E. WHITE 
Owing to the increased experience, the third classification given in 
Table 3 is probably the most accurate. Assuming that it is, from a 
total of 9,663 progeny, 6,999 were classed as white and 2,664 yellow. 
On the assumption of a one-factor difference between the two maize 
races with complete or practically complete dominance of the white 
color, the theoretically expected numbers would be 7,248 W : 2,416 Y 
(3 : i). The deviation between the ratio actually obtained and 
that theoretically expected is 249. The yellow segregates were far 
from uniform in color, all shades from a dark yellow (not orange) to 
a very light lemon yellow were present on the same ear. Further, in 
some ears, the yellow was principally confined to the base of the grain, 
nearest the point of attachment. A few dark yell'ow grains were 
somewhat deeper colored than the yellow grand-parental California 
Pop, but this may be due to segregation of various factors that effect 
endosperm texture, as the dark yellow grains usually appear less 
translucent than those of the grand-parental Pop variety. 
Back Crosses of Fi with Z 21 (the Dominant White Endosperm 
Parent) 
These two ears, resulting from back crosses of the Fi with the 
dominant endosperm parent, came from two Fi plants which also 
produced two selfed ears (296 and 34a in Table 3). The back-crossed 
ears were uniform in seed color, the white being more opaque than in 
the endosperm of the Fi grains. Nos. 2gb and 34a gave typical Fg 
ratios and the yellows were of several shades as in all the other ears 
with F2 seeds. 
From self-pollinated ears Nos. I, 2 and 3 (Table 3) of the F2 
generation, approximately 1,000 plants with F3 endosperm seeds were 
grown. Of these, 43 ears were self-pollinated, 27 of which came from 
F2 seeds classed as white, and 16 from F2 seeds classed as yellow. 
Nine of the white seeds gave all white F?, progeny, while 19 gave both 
white and yellow grains approximating the ratio of 3 W : i Y (Table 4). 
The 16 F2 seeds classed as yellows gave 14 all yellow ears, and 3 ears 
with both white and yellow seeds approximating the ratio of 3 W : i Y. 
(Z 14 X Z 2i)-29a X Z 21 
(Z 14 X Z 2i)-34b X Z 21 
All white 
All white 
F3 Generation 
