30 
ORLAND E. WHITE 
inferred that the parent plant (2-1A) had been largely homozygous in 
its genotypical constitution. 
From the cross referred to above (2-1 A X 321), 90 Fi progeny 
were grown, all of which were intermediate in both habit and in size 
of floral organs, but absolutely normal as regards pistillody. Two 
of these were selfed and F2 progeny grown. The results are tabulated 
in Table 4. 
Table 4 
Pedigree 
Normal 
Abnormal 
Total 
(394 X 321) — 2 — lA X 321 — I 
103 
82 
185 
(394 X 321) — 2 — lA X 321 — 2 
152 
42 
196 
Total 
257 
124 
381 
Expected 
285.75 
95-25 
381 
Deviation 
—28.75 
+28.75 
One family (-2) gave a fair approximation to the 3 :i ratio, but the other 
had a large excess of abnormal segregates, which I am at present unable 
to account for, because the two families were grown from the same 
grand parental stock, and under the same external environment. 
Many other characters of a structural nature had segregated in this F2 
generation, and the variation in the expression of the anomaly was 
large. Many plants were as abnormal, and many much less so than 
the grandmother. Other abnormalities appeared, both in pistilloid 
and normal segregates. Split corolla tubes and 3- to 4-loculed ovaries 
were not infrequent. Some of the segregates, as well as a number of 
the pure line (?) progeny, possessed flowers with pistilloid anthers 
containing numerous small ovules. Where these occurred, the pollen- 
sacs were deformed, sterile, and usually the ovules were exposed, owing 
to hypertrophy of the anther-sac walls. 
3. Catacorolla 
This is not an uncommon anomaly, and hereditary races of it have 
long been known, e. g., hose-in-hose primula, and a garden variety 
of gloxinia, first described by Prof. E. Morren (see Masters, 1869, 
pp. 451-52, figs. 213-14). Catacorolla has been exceedingly well 
described by both Morren and Masters, so I shall not take the space 
here for a general detailed description, but confine myself to the form 
it takes in the particular race with which I worked. This race (4-1 A) 
is descended from a single plant which possessed the catacorolla 
