UREDINALES OF GUATEMALA BASED ON COLLECTIONS 
BY E. W. D. HOLWAY 
L Introduction, Coleosporiaceae and Uredinaceae 
J. C. Arthur 
Guatemala, the largest of the Central American republics, and 
with much more than one third of their total population, is a land 
of great charm for the traveler and the naturalist. The hot, low lands 
near the coast, especially on the Atlantic side, with their dense tropical 
growth, the extensive plateau of the interior, ranging from 8,000 feet 
elevation in the north to half that altitude in the south, thus supplying 
a temperate climate, and the many high mountains with their pine- 
clad summits and cool breezes, give a wonderful range for all forms of 
vegetation. The large proportion of Indians among the population, 
the many cities of twenty-five to seventy-five thousand inhabitants, 
the diversity of landscape, and the enjoyable climate lend a special 
fascination to the task of the explorer. 
The rusts of Guatemala have been made known through the efforts 
of two tireless collectors of superior botanical attainments, who gave 
the Uredinales their first attention, endeavoring to take ample speci- 
mens to illustrate both the rust and its host, but who gathered also 
other fungi, as well as higher forms, especially phanerogams. Not a 
dozen collections of Guatemalan rusts are known from all other sources 
taken together. 
Professor W. A. Kellerman made four visits to Guatemala at the 
beginning of the years 1905, 1906, 1907 and 1908, and was so enamored 
of the country and its interesting vegetation that upon returning from 
his third trip in April, 1907, he laid plans for a peripatetic school of 
tropical botany. On his next visit he took with him a few students 
from the University of Ohio. The program for this visit had been 
completed and arrangements made for departure homeward when a 
brief illness terminated the career of a zealous and undaunted col- 
lector. The larger part of the rich material secured during these four 
years yet remains unstudied. Two papers dealing with the rust por- 
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