404 
HARRY M. FITZPATRICK 
to be four. Occasionally also four chromosome-like bodies can be 
seen in a nucleus which shows no indication of a spindle, and which 
is evidently in an early prophase (fig. 8). 
When the chromosomes begin to pass toward the poles the nuclear 
membrane is still evident, but it disappears soon afterward. In 
figure 15 the membrane is absent, and the chromosomes are shown 
scattered over the spindle. Here also the chromosome number is 
clearly shown to be four. The two spindles orient themselves parallel 
to the long axis of the cell, and soon come to lie side by side in typical 
conjugate division (fig. 16). The nucleolus is drawn into the spindle 
and passes toward one of the poles. In late telophase it is incorporated 
in one of the daughter nuclei. 
Although in metaphase and early anaphase the spindles do not 
occupy the position characteristic of conjugate division, they pass 
more or less definitely into this position before late telophase, and it is 
evident that the two daughter nuclei which migrate into each end of the 
cell are not in any case sisters. At the completion of the division the 
four resulting nuclei round up and remain for a brief period as well 
defined nuclei in a single cell (figs. 17, iS). The fact that these four- 
r^ucleate cells are rarely found indicates that the transverse septum 
is formed quickly. After the laying down of the septum the two re- 
sulting cells elongate, and the nuclei in each drift apart. 
As the sporophore approaches maturity, the hyphae at its periphery 
undergo a slight amount of branching, the terminal portions turning 
out at right angles to the long axis of the sporophore. A more or 
less definite palisade layer is thus formed. The terminal cells later 
undergo further growth, and develop into basidia. The basidia do 
not stand close together, and a definite hymenium, such as occurs in 
most of the higher Basidiomycetes, is not formed. Paraphyses are 
absent, and no sharp differentiation of the fruit-body into subhy- 
menium and trama occurs. The hyphae composing the sporophore 
interweave only to a slight degree, and a loose tissue results in which 
an individual hypha may be traced from the basidium far back into the 
fruit-body. 
Nuclear Phenomena in the Basidium 
When the basidium is merely the undifferentiated terminal cell 
of one of the hyphae composing the sporophore it contains two small 
nuclei similar to those present in other cells of the thread. As it 
