July, 1922] BAILEY — POLLINATION OF MARCGRAVIA 
377 
Concerning the identity of these Marcgravias, it may be stated at 
once that they both belong in Delpino's subgenus, Plagiothalamium. 
Furthermore, the prolongation of the peduncle beyond the points of attach- 
ment of the pedicels in species B indicates clearly that this species should 
be referred to Wittmack's subsection Dolichoracheae, just as the absence 
of this extension in species A leads to its inclusion in his Brachyracheae. 
The leaves and inflorescences of Marcgravias vary considerably, par- 
ticularly in different stages of their ontogeny, and are difficult to season 
for herbarium purposes. Many of the descriptions of species in the litera- 
ture appear to be based upon the study of somewhat fragmentary material; 
not infrequently of immature leaves or inflorescences. Such facts as these, 
coupled with the difficulty of securing access to the widely scattered type 
specimens, makes the determination of species a somewhat difffcult and 
unsatisfactory undertaking. 
Species B appears to be closely allied to M. coriacea Vahl and M. acu- 
minata Miguel, but differs from these species in certain of its floral and foliar 
characters, particularly in having filaments which are not basally connate. 
Species A resembles M. parviflora Rich. var. pedunculosa (Triana and 
Planchon) Wittm., but has large nectaries which are longer than the pedicels 
of the flowers. In view of the abundance of both living and dried material 
at my disposal, it seems wiser to describe these plants as new species, 
rather than to refer them to any of the above-named species. Further- 
more, it is desirable to make detailed and fairly comprehensive descriptions, 
since there appears to be some doubt as to which characters are of greatest 
specific significance in these sections of Marcgravia. 
Marcgravia (Dolichoracheae) cuyuniensis spec. nov. > 
Folia ramorum floralium breviter petiolata, oblongo-elliptica, coriacea, 
ex glandulis marginalibus depressis numerosis subcrenulata, apice 
acuminata, basi acuta vel subrotundata; in sicco haud nitentia, supra 
punctata, venis patentibus; glandulis hypophyllis saepissime duobus, ad 
basim costae mediae; nervo crasso subtus prominente, supra basim versus 
profunde sulcato; petiolis crassis antice canaliculatis. Racemus umbelli- 
formis, pendulus, submultiflorus, breviter pedunculatus. Bracteae 3-6, 
elongatae, tubuloso-cucullatae, crassae, clavatae, deciduae, sub ore ro- 
tundato plus minusve constrictae. Pedicelli 16-24, crassi, tuberculato- 
verrucosi. Corolla coriacea, ovoidea vel conico-ovoidea. Stamina zb 
46, filamentis basi non connatis, antheris rubellis. Ovarium 9- — 11- 
loculare. Fructus magnus, globosus, rubidus, plus minusve tuberculato- 
verrucosus, stigmate mammiformi coronatus. Semina sanguinea nitentia 
Scandent epiphytic shrub. Plagiotropic shoots quadrangular, bearing 
small coriaceous leaves and short clasping roots. The former oval, oval- 
oblong, or oblong, distichous, conspicuously glandular at the base. The 
latter aggregated in clusters, more or less completely sheltered by the 
leaves. Orthotropic fertile shoots numerous, short, stout, usually terete 
and pendulous, sparsely linear-verrucose. Leaves distichous, coriaceous, 
somewhat conduplicate dorsally, elliptical-oblong, acuminate, acute, retuse. 
